A1IGU5_Q96P47.1
contacts (residues)
Receptor
R: Secondary Structure
R: Residue
R; Chain
R: Residue number
Ligand
L: Secondary Structure
L: Residue
L: Chain
L: Residue Number
R
-
PRO
A
307
L
-
HIS
V
450
R
-
SER
A
308
L
-
HIS
V
449
R
-
SER
A
308
L
-
HIS
V
450
R
-
SER
A
308
L
-
VAL
V
451
R
-
SER
A
308
L
-
PRO
V
452
R
-
ILE
A
309
L
-
HIS
V
449
R
-
ILE
A
309
L
-
HIS
V
450
R
-
ILE
A
309
L
-
VAL
V
451
R
-
SER
A
310
L
H
MET
V
443
R
-
SER
A
310
L
-
HIS
V
449
R
-
GLN
A
311
L
H
MET
V
443
R
-
ARG
A
312
L
H
LEU
V
436
R
-
ARG
A
312
L
H
ALA
V
439
R
-
ARG
A
312
L
H
GLU
V
440
R
-
ARG
A
312
L
H
MET
V
443
R
-
GLU
A
313
L
H
LEU
V
436
R
-
LEU
A
314
L
H
ALA
V
432
R
-
LEU
A
314
L
H
VAL
V
435
R
-
LEU
A
314
L
H
LEU
V
436
R
-
ARG
A
315
L
H
ALA
V
432
R
-
ILE
A
316
L
H
VAL
V
425
R
-
ILE
A
316
L
H
GLN
V
428
R
-
ILE
A
316
L
H
ARG
V
429
R
-
ASN
A
447
L
H
GLU
V
112
R
-
GLY
A
448
L
H
GLU
V
112
R
-
LEU
A
449
L
H
LEU
V
108
R
-
LEU
A
449
L
H
GLN
V
111
R
-
LEU
A
449
L
H
GLU
V
112
R
-
SER
A
450
L
H
LEU
V
108
R
-
SER
A
450
L
H
GLN
V
111
R
-
SER
A
450
L
H
ARG
V
198
R
-
VAL
A
451
L
H
ASN
V
105
R
-
VAL
A
451
L
H
ILE
V
106
R
-
VAL
A
451
L
H
LEU
V
108
R
-
VAL
A
451
L
H
GLU
V
109
R
-
VAL
A
451
L
H
ARG
V
198
R
-
GLU
A
452
L
H
LEU
V
102
R
-
GLU
A
452
L
H
ASN
V
105
R
-
GLU
A
452
L
H
ILE
V
106
R
-
GLU
A
452
L
H
GLU
V
109
R
-
ARG
A
453
L
H
LEU
V
102
R
-
ARG
A
453
L
H
ASN
V
105
R
-
ARG
A
453
L
H
ILE
V
106
R
-
ARG
A
453
L
H
GLU
V
109
R
-
SER
A
454
L
H
LEU
V
102
R
-
SER
A
454
L
H
ILE
V
106
R
-
SER
A
454
L
H
ASP
V
88
R
-
SER
A
454
L
H
GLU
V
91
R
-
SER
A
454
L
H
THR
V
92
R
-
SER
A
454
L
-
LYS
V
95
R
-
ASN
A
455
L
H
ASP
V
88
R
-
ASN
A
455
L
H
LEU
V
89
R
-
ASN
A
455
L
H
GLN
V
90
R
-
ASN
A
455
L
H
GLU
V
91
R
-
ASN
A
455
L
H
THR
V
92
R
-
ASN
A
455
L
H
ALA
V
93
R
-
ASN
A
455
L
H
SER
V
94
R
-
ASN
A
455
L
-
LYS
V
95
R
-
THR
A
456
L
H
LEU
V
89
R
-
THR
A
456
L
H
GLN
V
90
R
-
THR
A
456
L
H
GLU
V
91
R
-
THR
A
456
L
H
THR
V
92
R
-
THR
A
456
L
H
ALA
V
93
R
-
THR
A
456
L
H
SER
V
94
R
-
THR
A
456
L
-
LYS
V
95
R
-
GLN
A
457
L
H
ILE
V
39
R
-
GLN
A
457
L
H
GLN
V
90
R
-
GLN
A
457
L
H
GLU
V
91
R
-
GLN
A
457
L
H
ALA
V
93
R
-
LEU
A
458
L
H
ILE
V
39
R
-
LEU
A
458
L
H
ASP
V
40
R
-
LEU
A
458
L
H
GLN
V
90
R
-
GLY
A
459
L
H
ARG
V
36
R
-
GLY
A
459
L
H
ILE
V
39
R
-
GLY
A
459
L
H
ASP
V
40
R
-
GLY
A
459
L
H
VAL
V
43
R
-
GLY
A
460
L
H
ASP
V
40
R
-
GLY
A
460
L
H
VAL
V
43
R
-
GLY
A
461
L
H
ASP
V
40
R
-
GLY
A
461
L
H
THR
V
41
R
-
GLY
A
461
L
H
SER
V
44
R
-
THR
A
462
L
H
SER
V
44
R
-
GLY
A
463
L
H
ARG
V
174
R
-
ALA
A
464
L
H
PHE
V
163
R
-
ALA
A
464
L
H
ARG
V
174
R
-
PRO
A
465
L
H
PHE
V
163
R
-
PRO
A
465
L
H
GLN
V
170
R
-
PRO
A
465
L
H
ARG
V
174
R
-
HIS
A
466
L
-
VAL
V
166
R
-
HIS
A
466
L
H
GLN
V
170
R
-
HIS
A
466
L
H
ARG
V
174
R
-
SER
A
467
L
-
VAL
V
166
R
-
SER
A
467
L
H
GLN
V
170
R
-
ALA
A
468
L
H
TYR
V
125
R
-
ALA
A
468
L
-
VAL
V
166
R
-
ALA
A
468
L
H
LEU
V
169
R
-
ALA
A
468
L
H
GLN
V
170
R
-
SER
A
469
L
H
TYR
V
125
R
-
SER
A
469
L
H
LEU
V
129
R
-
SER
A
470
L
H
LEU
V
129
R
-
ALA
A
471
L
H
LEU
V
129
R
-
ALA
A
471
L
H
LEU
V
130
R
-
ALA
A
471
L
H
ASP
V
133
R
-
SER
A
472
L
H
LEU
V
130
R
-
SER
A
472
L
H
ASP
V
133
R
-
LEU
A
473
L
H
LYS
V
137
R
-
HIS
A
474
L
H
LYS
V
137
R
-
SER
A
475
L
H
LYS
V
137
R
-
GLU
A
476
L
H
LYS
V
137
R
-
ARG
A
477
L
H
PRO
V
139
R
-
PRO
A
478
L
H
PRO
V
139
R
-
LEU
A
479
L
H
PRO
V
139
R
-
LEU
A
479
L
H
ARG
V
143
R
-
SER
A
480
L
H
ARG
V
143
R
H
LEU
A
620
L
H
ARG
V
429
R
H
GLN
A
623
L
H
GLN
V
419
R
H
ASN
A
624
L
H
LEU
V
342
R
-
ARG
A
636
L
-
PRO
V
64
R
-
GLY
A
637
L
H
GLN
V
65
R
-
PHE
A
640
L
H
GLN
V
65
A1IGU5_Q96P47.1
contacts (residues & atoms)
Receptor
R: Secondary Structure
R: Atom
R: Residue
R; Chain
R: Residue number
Ligand
L: Secondary Structure
L; Atom
L: Residue
L: Chain
L: Residue Number
R
-
N
PRO
A
307
L
-
HB3
HIS
V
450
R
-
CA
PRO
A
307
L
-
HB3
HIS
V
450
R
-
C
PRO
A
307
L
-
CA
HIS
V
450
R
-
C
PRO
A
307
L
-
CB
HIS
V
450
R
-
C
PRO
A
307
L
-
O
HIS
V
450
R
-
C
PRO
A
307
L
-
ND1
HIS
V
450
R
-
C
PRO
A
307
L
-
HA
HIS
V
450
R
-
C
PRO
A
307
L
-
HB3
HIS
V
450
R
-
CB
PRO
A
307
L
-
O
HIS
V
450
R
-
CB
PRO
A
307
L
-
HB3
HIS
V
450
R
-
O
PRO
A
307
L
-
CB
HIS
V
450
R
-
O
PRO
A
307
L
-
CG
HIS
V
450
R
-
O
PRO
A
307
L
-
ND1
HIS
V
450
R
-
O
PRO
A
307
L
-
CE1
HIS
V
450
R
-
O
PRO
A
307
L
-
HA
HIS
V
450
R
-
O
PRO
A
307
L
-
HB3
HIS
V
450
R
-
CG
PRO
A
307
L
-
O
HIS
V
450
R
-
CG
PRO
A
307
L
-
HB3
HIS
V
450
R
-
CD
PRO
A
307
L
-
HB3
HIS
V
450
R
-
N
SER
A
308
L
-
CA
HIS
V
450
R
-
N
SER
A
308
L
-
C
HIS
V
450
R
-
N
SER
A
308
L
-
CB
HIS
V
450
R
-
N
SER
A
308
L
-
O
HIS
V
450
R
-
N
SER
A
308
L
-
HA
HIS
V
450
R
-
N
SER
A
308
L
-
HB3
HIS
V
450
R
-
CA
SER
A
308
L
-
CA
HIS
V
450
R
-
CA
SER
A
308
L
-
O
HIS
V
450
R
-
CA
SER
A
308
L
-
HA
HIS
V
450
R
-
C
SER
A
308
L
-
O
HIS
V
449
R
-
C
SER
A
308
L
-
CA
HIS
V
450
R
-
C
SER
A
308
L
-
C
HIS
V
450
R
-
C
SER
A
308
L
-
O
HIS
V
450
R
-
C
SER
A
308
L
-
HA
HIS
V
450
R
-
O
SER
A
308
L
-
O
HIS
V
449
R
-
O
SER
A
308
L
-
CA
HIS
V
450
R
-
O
SER
A
308
L
-
C
HIS
V
450
R
-
O
SER
A
308
L
-
O
HIS
V
450
R
-
O
SER
A
308
L
-
HA
HIS
V
450
R
-
O
SER
A
308
L
-
N
VAL
V
451
R
-
O
SER
A
308
L
-
C
VAL
V
451
R
-
O
SER
A
308
L
-
O
VAL
V
451
R
-
O
SER
A
308
L
-
N
PRO
V
452
R
-
O
SER
A
308
L
-
HA
PRO
V
452
R
-
N
ILE
A
309
L
-
O
HIS
V
449
R
-
N
ILE
A
309
L
-
HA
HIS
V
450
R
-
CA
ILE
A
309
L
-
O
HIS
V
449
R
-
C
ILE
A
309
L
-
C
HIS
V
449
R
-
C
ILE
A
309
L
-
O
HIS
V
449
R
-
C
ILE
A
309
L
-
HA
HIS
V
449
R
-
O
ILE
A
309
L
-
C
HIS
V
449
R
-
O
ILE
A
309
L
-
O
HIS
V
449
R
-
O
ILE
A
309
L
-
HA
HIS
V
449
R
-
O
ILE
A
309
L
-
O
VAL
V
451
R
-
N
SER
A
310
L
-
O
HIS
V
449
R
-
N
SER
A
310
L
-
CD2
HIS
V
449
R
-
N
SER
A
310
L
-
NE2
HIS
V
449
R
-
N
SER
A
310
L
-
HD2
HIS
V
449
R
-
N
SER
A
310
L
-
HE2
HIS
V
449
R
-
C
SER
A
310
L
-
HD2
HIS
V
449
R
-
O
SER
A
310
L
H
CB
MET
V
443
R
-
O
SER
A
310
L
H
SD
MET
V
443
R
-
O
SER
A
310
L
H
HB2
MET
V
443
R
-
O
SER
A
310
L
H
HE3
MET
V
443
R
-
O
SER
A
310
L
-
CD2
HIS
V
449
R
-
O
SER
A
310
L
-
NE2
HIS
V
449
R
-
O
SER
A
310
L
-
HD2
HIS
V
449
R
-
O
SER
A
310
L
-
HE2
HIS
V
449
R
-
OG
SER
A
310
L
-
HA
HIS
V
449
R
-
N
GLN
A
311
L
H
CE
MET
V
443
R
-
N
GLN
A
311
L
H
HE1
MET
V
443
R
-
N
GLN
A
311
L
H
HE3
MET
V
443
R
-
CA
GLN
A
311
L
H
HE1
MET
V
443
R
-
C
GLN
A
311
L
H
SD
MET
V
443
R
-
C
GLN
A
311
L
H
CE
MET
V
443
R
-
C
GLN
A
311
L
H
HE1
MET
V
443
R
-
O
GLN
A
311
L
H
SD
MET
V
443
R
-
O
GLN
A
311
L
H
CE
MET
V
443
R
-
O
GLN
A
311
L
H
HE1
MET
V
443
R
-
N
ARG
A
312
L
H
SD
MET
V
443
R
-
N
ARG
A
312
L
H
CE
MET
V
443
R
-
N
ARG
A
312
L
H
HE1
MET
V
443
R
-
N
ARG
A
312
L
H
HE2
MET
V
443
R
-
N
ARG
A
312
L
H
HE3
MET
V
443
R
-
CA
ARG
A
312
L
H
HE1
MET
V
443
R
-
C
ARG
A
312
L
H
CD1
LEU
V
436
R
-
C
ARG
A
312
L
H
HD11
LEU
V
436
R
-
C
ARG
A
312
L
H
HD12
LEU
V
436
R
-
C
ARG
A
312
L
H
HD13
LEU
V
436
R
-
C
ARG
A
312
L
H
HE1
MET
V
443
R
-
CB
ARG
A
312
L
H
HE1
MET
V
443
R
-
O
ARG
A
312
L
H
CD1
LEU
V
436
R
-
O
ARG
A
312
L
H
HA
LEU
V
436
R
-
O
ARG
A
312
L
H
HD11
LEU
V
436
R
-
O
ARG
A
312
L
H
HD12
LEU
V
436
R
-
O
ARG
A
312
L
H
HD13
LEU
V
436
R
-
O
ARG
A
312
L
H
HB1
ALA
V
439
R
-
O
ARG
A
312
L
H
HB2
ALA
V
439
R
-
O
ARG
A
312
L
H
HG2
GLU
V
440
R
-
O
ARG
A
312
L
H
CE
MET
V
443
R
-
O
ARG
A
312
L
H
HE1
MET
V
443
R
-
O
ARG
A
312
L
H
HE2
MET
V
443
R
-
N
GLU
A
313
L
H
CD1
LEU
V
436
R
-
N
GLU
A
313
L
H
HD11
LEU
V
436
R
-
N
GLU
A
313
L
H
HD12
LEU
V
436
R
-
N
GLU
A
313
L
H
HD13
LEU
V
436
R
-
CA
GLU
A
313
L
H
CD1
LEU
V
436
R
-
CA
GLU
A
313
L
H
HD11
LEU
V
436
R
-
CA
GLU
A
313
L
H
HD12
LEU
V
436
R
-
CA
GLU
A
313
L
H
HD13
LEU
V
436
R
-
C
GLU
A
313
L
H
CD1
LEU
V
436
R
-
C
GLU
A
313
L
H
HD12
LEU
V
436
R
-
O
GLU
A
313
L
H
HD12
LEU
V
436
R
-
N
LEU
A
314
L
H
HB2
LEU
V
436
R
-
N
LEU
A
314
L
H
HD12
LEU
V
436
R
-
N
LEU
A
314
L
H
HD23
LEU
V
436
R
-
CA
LEU
A
314
L
H
HB1
ALA
V
432
R
-
C
LEU
A
314
L
H
HB1
ALA
V
432
R
-
CB
LEU
A
314
L
H
CB
ALA
V
432
R
-
CB
LEU
A
314
L
H
O
ALA
V
432
R
-
CB
LEU
A
314
L
H
HA
ALA
V
432
R
-
CB
LEU
A
314
L
H
HB1
ALA
V
432
R
-
CB
LEU
A
314
L
H
HB2
LEU
V
436
R
-
O
LEU
A
314
L
H
HB1
ALA
V
432
R
-
O
LEU
A
314
L
H
HD23
LEU
V
436
R
-
CG
LEU
A
314
L
H
HA
ALA
V
432
R
-
CG
LEU
A
314
L
H
HB1
ALA
V
432
R
-
CD1
LEU
A
314
L
H
HA
ALA
V
432
R
-
CD1
LEU
A
314
L
H
CB
VAL
V
435
R
-
CD1
LEU
A
314
L
H
CG1
VAL
V
435
R
-
CD1
LEU
A
314
L
H
HB
VAL
V
435
R
-
CD1
LEU
A
314
L
H
HG11
VAL
V
435
R
-
CD1
LEU
A
314
L
H
HG13
VAL
V
435
R
-
CD1
LEU
A
314
L
H
HG22
VAL
V
435
R
-
N
ARG
A
315
L
H
HB1
ALA
V
432
R
-
CB
ILE
A
316
L
H
HA
ARG
V
429
R
-
CB
ILE
A
316
L
H
HB2
ARG
V
429
R
-
CB
ILE
A
316
L
H
HG3
ARG
V
429
R
-
CG1
ILE
A
316
L
H
N
ARG
V
429
R
-
CG1
ILE
A
316
L
H
CA
ARG
V
429
R
-
CG1
ILE
A
316
L
H
CB
ARG
V
429
R
-
CG1
ILE
A
316
L
H
HA
ARG
V
429
R
-
CG1
ILE
A
316
L
H
HB2
ARG
V
429
R
-
CG1
ILE
A
316
L
H
HG3
ARG
V
429
R
-
CG2
ILE
A
316
L
H
HG13
VAL
V
425
R
-
CG2
ILE
A
316
L
H
HB2
ARG
V
429
R
-
CD1
ILE
A
316
L
H
O
VAL
V
425
R
-
CD1
ILE
A
316
L
H
C
GLN
V
428
R
-
CD1
ILE
A
316
L
H
CB
GLN
V
428
R
-
CD1
ILE
A
316
L
H
O
GLN
V
428
R
-
CD1
ILE
A
316
L
H
HB2
GLN
V
428
R
-
CD1
ILE
A
316
L
H
HB3
GLN
V
428
R
-
CD1
ILE
A
316
L
H
N
ARG
V
429
R
-
CD1
ILE
A
316
L
H
CA
ARG
V
429
R
-
CD1
ILE
A
316
L
H
C
ARG
V
429
R
-
CD1
ILE
A
316
L
H
CB
ARG
V
429
R
-
CD1
ILE
A
316
L
H
CG
ARG
V
429
R
-
CD1
ILE
A
316
L
H
H
ARG
V
429
R
-
CD1
ILE
A
316
L
H
HA
ARG
V
429
R
-
CD1
ILE
A
316
L
H
HB2
ARG
V
429
R
-
CD1
ILE
A
316
L
H
HB3
ARG
V
429
R
-
CD1
ILE
A
316
L
H
HG3
ARG
V
429
R
-
C
ASN
A
447
L
H
OE1
GLU
V
112
R
-
O
ASN
A
447
L
H
CD
GLU
V
112
R
-
O
ASN
A
447
L
H
OE1
GLU
V
112
R
-
N
GLY
A
448
L
H
OE1
GLU
V
112
R
-
CA
GLY
A
448
L
H
OE1
GLU
V
112
R
-
C
GLY
A
448
L
H
OE1
GLU
V
112
R
-
O
GLY
A
448
L
H
OE1
GLU
V
112
R
-
N
LEU
A
449
L
H
NE2
GLN
V
111
R
-
N
LEU
A
449
L
H
HE22
GLN
V
111
R
-
N
LEU
A
449
L
H
OE1
GLU
V
112
R
-
CA
LEU
A
449
L
H
HE22
GLN
V
111
R
-
C
LEU
A
449
L
H
NE2
GLN
V
111
R
-
C
LEU
A
449
L
H
HE21
GLN
V
111
R
-
C
LEU
A
449
L
H
HE22
GLN
V
111
R
-
CB
LEU
A
449
L
H
HE22
GLN
V
111
R
-
O
LEU
A
449
L
H
O
LEU
V
108
R
-
O
LEU
A
449
L
H
NE2
GLN
V
111
R
-
O
LEU
A
449
L
H
HG2
GLN
V
111
R
-
O
LEU
A
449
L
H
HE21
GLN
V
111
R
-
O
LEU
A
449
L
H
HE22
GLN
V
111
R
-
CA
SER
A
450
L
H
HB3
LEU
V
108
R
-
C
SER
A
450
L
H
CB
LEU
V
108
R
-
C
SER
A
450
L
H
HB2
LEU
V
108
R
-
C
SER
A
450
L
H
HB3
LEU
V
108
R
-
C
SER
A
450
L
H
HD12
LEU
V
108
R
-
O
SER
A
450
L
H
CB
LEU
V
108
R
-
O
SER
A
450
L
H
CG
LEU
V
108
R
-
O
SER
A
450
L
H
CD1
LEU
V
108
R
-
O
SER
A
450
L
H
CD2
LEU
V
108
R
-
O
SER
A
450
L
H
HB2
LEU
V
108
R
-
O
SER
A
450
L
H
HB3
LEU
V
108
R
-
O
SER
A
450
L
H
HD12
LEU
V
108
R
-
O
SER
A
450
L
H
HD22
LEU
V
108
R
-
O
SER
A
450
L
H
HE21
GLN
V
111
R
-
O
SER
A
450
L
H
HE22
GLN
V
111
R
-
O
SER
A
450
L
H
HH12
ARG
V
198
R
-
N
VAL
A
451
L
H
C
LEU
V
108
R
-
N
VAL
A
451
L
H
CB
LEU
V
108
R
-
N
VAL
A
451
L
H
HB2
LEU
V
108
R
-
N
VAL
A
451
L
H
HB3
LEU
V
108
R
-
N
VAL
A
451
L
H
HD11
LEU
V
108
R
-
N
VAL
A
451
L
H
HD12
LEU
V
108
R
-
N
VAL
A
451
L
H
N
GLU
V
109
R
-
CA
VAL
A
451
L
H
HA
ASN
V
105
R
-
CA
VAL
A
451
L
H
HB3
ASN
V
105
R
-
CA
VAL
A
451
L
H
HB2
LEU
V
108
R
-
CA
VAL
A
451
L
H
HB3
LEU
V
108
R
-
C
VAL
A
451
L
H
CA
ASN
V
105
R
-
C
VAL
A
451
L
H
C
ASN
V
105
R
-
C
VAL
A
451
L
H
CB
ASN
V
105
R
-
C
VAL
A
451
L
H
O
ASN
V
105
R
-
C
VAL
A
451
L
H
CG
ASN
V
105
R
-
C
VAL
A
451
L
H
OD1
ASN
V
105
R
-
C
VAL
A
451
L
H
HA
ASN
V
105
R
-
C
VAL
A
451
L
H
HB3
ASN
V
105
R
-
C
VAL
A
451
L
H
HB2
LEU
V
108
R
-
C
VAL
A
451
L
H
HB3
GLU
V
109
R
-
CB
VAL
A
451
L
H
OD1
ASN
V
105
R
-
CB
VAL
A
451
L
H
HA
ASN
V
105
R
-
CB
VAL
A
451
L
H
HD11
LEU
V
108
R
-
CB
VAL
A
451
L
H
HD12
LEU
V
108
R
-
CB
VAL
A
451
L
H
HH12
ARG
V
198
R
-
CB
VAL
A
451
L
H
HH22
ARG
V
198
R
-
O
VAL
A
451
L
H
CA
ASN
V
105
R
-
O
VAL
A
451
L
H
C
ASN
V
105
R
-
O
VAL
A
451
L
H
CB
ASN
V
105
R
-
O
VAL
A
451
L
H
O
ASN
V
105
R
-
O
VAL
A
451
L
H
CG
ASN
V
105
R
-
O
VAL
A
451
L
H
HA
ASN
V
105
R
-
O
VAL
A
451
L
H
HB2
ASN
V
105
R
-
O
VAL
A
451
L
H
HB3
ASN
V
105
R
-
O
VAL
A
451
L
H
N
ILE
V
106
R
-
O
VAL
A
451
L
H
CB
LEU
V
108
R
-
O
VAL
A
451
L
H
HB2
LEU
V
108
R
-
O
VAL
A
451
L
H
HB3
LEU
V
108
R
-
O
VAL
A
451
L
H
N
GLU
V
109
R
-
O
VAL
A
451
L
H
CB
GLU
V
109
R
-
O
VAL
A
451
L
H
H
GLU
V
109
R
-
O
VAL
A
451
L
H
HB3
GLU
V
109
R
-
CG1
VAL
A
451
L
H
CG
ASN
V
105
R
-
CG1
VAL
A
451
L
H
ND2
ASN
V
105
R
-
CG1
VAL
A
451
L
H
OD1
ASN
V
105
R
-
CG1
VAL
A
451
L
H
HB3
ASN
V
105
R
-
CG1
VAL
A
451
L
H
HD22
ASN
V
105
R
-
CG1
VAL
A
451
L
H
HH22
ARG
V
198
R
-
CG2
VAL
A
451
L
H
NH1
ARG
V
198
R
-
CG2
VAL
A
451
L
H
HH12
ARG
V
198
R
-
CG2
VAL
A
451
L
H
HH22
ARG
V
198
R
-
N
GLU
A
452
L
H
CB
ASN
V
105
R
-
N
GLU
A
452
L
H
CG
ASN
V
105
R
-
N
GLU
A
452
L
H
HB3
ASN
V
105
R
-
N
GLU
A
452
L
H
HB3
GLU
V
109
R
-
CA
GLU
A
452
L
H
CB
ASN
V
105
R
-
CA
GLU
A
452
L
H
CG
ASN
V
105
R
-
CA
GLU
A
452
L
H
HB2
ASN
V
105
R
-
CA
GLU
A
452
L
H
HB3
ASN
V
105
R
-
CA
GLU
A
452
L
H
HD21
ASN
V
105
R
-
C
GLU
A
452
L
H
CA
ASN
V
105
R
-
C
GLU
A
452
L
H
C
ASN
V
105
R
-
C
GLU
A
452
L
H
CB
ASN
V
105
R
-
C
GLU
A
452
L
H
O
ASN
V
105
R
-
C
GLU
A
452
L
H
CG
ASN
V
105
R
-
C
GLU
A
452
L
H
HB2
ASN
V
105
R
-
C
GLU
A
452
L
H
HB3
ASN
V
105
R
-
C
GLU
A
452
L
H
HD21
ASN
V
105
R
-
C
GLU
A
452
L
H
N
ILE
V
106
R
-
C
GLU
A
452
L
H
H
ILE
V
106
R
-
C
GLU
A
452
L
H
HA
ILE
V
106
R
-
C
GLU
A
452
L
H
HG13
ILE
V
106
R
-
C
GLU
A
452
L
H
OE2
GLU
V
109
R
-
CB
GLU
A
452
L
H
CB
ASN
V
105
R
-
CB
GLU
A
452
L
H
CG
ASN
V
105
R
-
CB
GLU
A
452
L
H
ND2
ASN
V
105
R
-
CB
GLU
A
452
L
H
HB2
ASN
V
105
R
-
CB
GLU
A
452
L
H
HB3
ASN
V
105
R
-
CB
GLU
A
452
L
H
HD21
ASN
V
105
R
-
CB
GLU
A
452
L
H
HD22
ASN
V
105
R
-
O
GLU
A
452
L
H
O
LEU
V
102
R
-
O
GLU
A
452
L
H
HA
LEU
V
102
R
-
O
GLU
A
452
L
H
N
ASN
V
105
R
-
O
GLU
A
452
L
H
CA
ASN
V
105
R
-
O
GLU
A
452
L
H
C
ASN
V
105
R
-
O
GLU
A
452
L
H
CB
ASN
V
105
R
-
O
GLU
A
452
L
H
O
ASN
V
105
R
-
O
GLU
A
452
L
H
CG
ASN
V
105
R
-
O
GLU
A
452
L
H
ND2
ASN
V
105
R
-
O
GLU
A
452
L
H
OD1
ASN
V
105
R
-
O
GLU
A
452
L
H
H
ASN
V
105
R
-
O
GLU
A
452
L
H
HA
ASN
V
105
R
-
O
GLU
A
452
L
H
HB2
ASN
V
105
R
-
O
GLU
A
452
L
H
HB3
ASN
V
105
R
-
O
GLU
A
452
L
H
HD21
ASN
V
105
R
-
O
GLU
A
452
L
H
N
ILE
V
106
R
-
O
GLU
A
452
L
H
CA
ILE
V
106
R
-
O
GLU
A
452
L
H
H
ILE
V
106
R
-
O
GLU
A
452
L
H
HA
ILE
V
106
R
-
O
GLU
A
452
L
H
HG13
ILE
V
106
R
-
N
ARG
A
453
L
H
CB
ASN
V
105
R
-
N
ARG
A
453
L
H
HB2
ASN
V
105
R
-
N
ARG
A
453
L
H
HB3
ASN
V
105
R
-
N
ARG
A
453
L
H
N
ILE
V
106
R
-
N
ARG
A
453
L
H
CG1
ILE
V
106
R
-
N
ARG
A
453
L
H
HA
ILE
V
106
R
-
N
ARG
A
453
L
H
HG12
ILE
V
106
R
-
N
ARG
A
453
L
H
HG13
ILE
V
106
R
-
N
ARG
A
453
L
H
OE2
GLU
V
109
R
-
CA
ARG
A
453
L
H
HD23
LEU
V
102
R
-
CA
ARG
A
453
L
H
HG12
ILE
V
106
R
-
CA
ARG
A
453
L
H
HG13
ILE
V
106
R
-
CA
ARG
A
453
L
H
OE2
GLU
V
109
R
-
C
ARG
A
453
L
H
CD2
LEU
V
102
R
-
C
ARG
A
453
L
H
HB3
LEU
V
102
R
-
C
ARG
A
453
L
H
HD21
LEU
V
102
R
-
C
ARG
A
453
L
H
HD23
LEU
V
102
R
-
C
ARG
A
453
L
H
HG13
ILE
V
106
R
-
O
ARG
A
453
L
H
CG
LEU
V
102
R
-
O
ARG
A
453
L
H
CD2
LEU
V
102
R
-
O
ARG
A
453
L
H
HB3
LEU
V
102
R
-
O
ARG
A
453
L
H
HD21
LEU
V
102
R
-
O
ARG
A
453
L
H
HD22
LEU
V
102
R
-
O
ARG
A
453
L
H
HD23
LEU
V
102
R
-
N
SER
A
454
L
H
HG23
THR
V
92
R
-
N
SER
A
454
L
H
CD2
LEU
V
102
R
-
N
SER
A
454
L
H
HB3
LEU
V
102
R
-
N
SER
A
454
L
H
HD12
LEU
V
102
R
-
N
SER
A
454
L
H
HD23
LEU
V
102
R
-
N
SER
A
454
L
H
HG13
ILE
V
106
R
-
N
SER
A
454
L
H
HD11
ILE
V
106
R
-
CA
SER
A
454
L
H
HG23
THR
V
92
R
-
CA
SER
A
454
L
-
HZ3
LYS
V
95
R
-
CA
SER
A
454
L
H
HB3
LEU
V
102
R
-
CA
SER
A
454
L
H
HD12
LEU
V
102
R
-
CA
SER
A
454
L
H
HD23
LEU
V
102
R
-
CA
SER
A
454
L
H
HD11
ILE
V
106
R
-
C
SER
A
454
L
H
O
ASP
V
88
R
-
C
SER
A
454
L
H
C
GLU
V
91
R
-
C
SER
A
454
L
H
CB
GLU
V
91
R
-
C
SER
A
454
L
H
HB2
GLU
V
91
R
-
C
SER
A
454
L
H
HB3
GLU
V
91
R
-
C
SER
A
454
L
H
N
THR
V
92
R
-
C
SER
A
454
L
H
CA
THR
V
92
R
-
C
SER
A
454
L
H
CG2
THR
V
92
R
-
C
SER
A
454
L
H
H
THR
V
92
R
-
C
SER
A
454
L
H
HA
THR
V
92
R
-
C
SER
A
454
L
H
HG22
THR
V
92
R
-
C
SER
A
454
L
H
HG23
THR
V
92
R
-
C
SER
A
454
L
-
HZ3
LYS
V
95
R
-
C
SER
A
454
L
H
HB3
LEU
V
102
R
-
C
SER
A
454
L
H
HD12
LEU
V
102
R
-
C
SER
A
454
L
H
HD23
LEU
V
102
R
-
C
SER
A
454
L
H
HD11
ILE
V
106
R
-
CB
SER
A
454
L
-
CE
LYS
V
95
R
-
CB
SER
A
454
L
-
NZ
LYS
V
95
R
-
CB
SER
A
454
L
-
HD2
LYS
V
95
R
-
CB
SER
A
454
L
-
HZ1
LYS
V
95
R
-
CB
SER
A
454
L
-
HZ2
LYS
V
95
R
-
CB
SER
A
454
L
-
HZ3
LYS
V
95
R
-
CB
SER
A
454
L
H
CD1
LEU
V
102
R
-
CB
SER
A
454
L
H
CD2
LEU
V
102
R
-
CB
SER
A
454
L
H
HD12
LEU
V
102
R
-
CB
SER
A
454
L
H
HD13
LEU
V
102
R
-
CB
SER
A
454
L
H
HD22
LEU
V
102
R
-
CB
SER
A
454
L
H
HD23
LEU
V
102
R
-
O
SER
A
454
L
H
O
ASP
V
88
R
-
O
SER
A
454
L
H
C
GLU
V
91
R
-
O
SER
A
454
L
H
N
THR
V
92
R
-
O
SER
A
454
L
H
CA
THR
V
92
R
-
O
SER
A
454
L
H
CB
THR
V
92
R
-
O
SER
A
454
L
H
CG2
THR
V
92
R
-
O
SER
A
454
L
H
H
THR
V
92
R
-
O
SER
A
454
L
H
HA
THR
V
92
R
-
O
SER
A
454
L
H
HB
THR
V
92
R
-
O
SER
A
454
L
H
HG21
THR
V
92
R
-
O
SER
A
454
L
H
HG22
THR
V
92
R
-
O
SER
A
454
L
H
HG23
THR
V
92
R
-
O
SER
A
454
L
-
HD2
LYS
V
95
R
-
O
SER
A
454
L
H
CB
LEU
V
102
R
-
O
SER
A
454
L
H
CG
LEU
V
102
R
-
O
SER
A
454
L
H
CD1
LEU
V
102
R
-
O
SER
A
454
L
H
HB2
LEU
V
102
R
-
O
SER
A
454
L
H
HB3
LEU
V
102
R
-
O
SER
A
454
L
H
HD11
LEU
V
102
R
-
O
SER
A
454
L
H
HD12
LEU
V
102
R
-
O
SER
A
454
L
H
HD13
LEU
V
102
R
-
O
SER
A
454
L
H
HD23
LEU
V
102
R
-
O
SER
A
454
L
H
HD11
ILE
V
106
R
-
OG
SER
A
454
L
-
NZ
LYS
V
95
R
-
OG
SER
A
454
L
-
HZ1
LYS
V
95
R
-
OG
SER
A
454
L
-
HZ2
LYS
V
95
R
-
OG
SER
A
454
L
-
HZ3
LYS
V
95
R
-
OG
SER
A
454
L
H
CD2
LEU
V
102
R
-
OG
SER
A
454
L
H
HD12
LEU
V
102
R
-
OG
SER
A
454
L
H
HD22
LEU
V
102
R
-
OG
SER
A
454
L
H
HD23
LEU
V
102
R
-
N
ASN
A
455
L
H
O
ASP
V
88
R
-
N
ASN
A
455
L
H
CA
GLU
V
91
R
-
N
ASN
A
455
L
H
C
GLU
V
91
R
-
N
ASN
A
455
L
H
CB
GLU
V
91
R
-
N
ASN
A
455
L
H
O
GLU
V
91
R
-
N
ASN
A
455
L
H
CG
GLU
V
91
R
-
N
ASN
A
455
L
H
OE2
GLU
V
91
R
-
N
ASN
A
455
L
H
HB2
GLU
V
91
R
-
N
ASN
A
455
L
H
HB3
GLU
V
91
R
-
N
ASN
A
455
L
H
N
THR
V
92
R
-
N
ASN
A
455
L
H
CA
THR
V
92
R
-
N
ASN
A
455
L
H
H
THR
V
92
R
-
N
ASN
A
455
L
H
HA
THR
V
92
R
-
N
ASN
A
455
L
H
HG22
THR
V
92
R
-
N
ASN
A
455
L
H
HG23
THR
V
92
R
-
CA
ASN
A
455
L
H
O
ASP
V
88
R
-
CA
ASN
A
455
L
H
N
GLU
V
91
R
-
CA
ASN
A
455
L
H
CA
GLU
V
91
R
-
CA
ASN
A
455
L
H
C
GLU
V
91
R
-
CA
ASN
A
455
L
H
CB
GLU
V
91
R
-
CA
ASN
A
455
L
H
O
GLU
V
91
R
-
CA
ASN
A
455
L
H
CG
GLU
V
91
R
-
CA
ASN
A
455
L
H
CD
GLU
V
91
R
-
CA
ASN
A
455
L
H
H
GLU
V
91
R
-
CA
ASN
A
455
L
H
HA
GLU
V
91
R
-
CA
ASN
A
455
L
H
HB2
GLU
V
91
R
-
CA
ASN
A
455
L
H
HB3
GLU
V
91
R
-
CA
ASN
A
455
L
H
HG2
GLU
V
91
R
-
CA
ASN
A
455
L
H
N
THR
V
92
R
-
CA
ASN
A
455
L
H
CA
THR
V
92
R
-
CA
ASN
A
455
L
H
H
THR
V
92
R
-
CA
ASN
A
455
L
H
HA
THR
V
92
R
-
CA
ASN
A
455
L
H
HG
SER
V
94
R
-
C
ASN
A
455
L
H
O
ASP
V
88
R
-
C
ASN
A
455
L
H
C
GLN
V
90
R
-
C
ASN
A
455
L
H
O
GLN
V
90
R
-
C
ASN
A
455
L
H
N
GLU
V
91
R
-
C
ASN
A
455
L
H
CA
GLU
V
91
R
-
C
ASN
A
455
L
H
C
GLU
V
91
R
-
C
ASN
A
455
L
H
CB
GLU
V
91
R
-
C
ASN
A
455
L
H
O
GLU
V
91
R
-
C
ASN
A
455
L
H
CG
GLU
V
91
R
-
C
ASN
A
455
L
H
H
GLU
V
91
R
-
C
ASN
A
455
L
H
HA
GLU
V
91
R
-
C
ASN
A
455
L
H
HB2
GLU
V
91
R
-
C
ASN
A
455
L
H
HB3
GLU
V
91
R
-
C
ASN
A
455
L
H
HG2
GLU
V
91
R
-
C
ASN
A
455
L
H
N
THR
V
92
R
-
C
ASN
A
455
L
H
CA
THR
V
92
R
-
C
ASN
A
455
L
H
C
THR
V
92
R
-
C
ASN
A
455
L
H
CB
THR
V
92
R
-
C
ASN
A
455
L
H
CG2
THR
V
92
R
-
C
ASN
A
455
L
H
H
THR
V
92
R
-
C
ASN
A
455
L
H
HA
THR
V
92
R
-
C
ASN
A
455
L
H
HG22
THR
V
92
R
-
C
ASN
A
455
L
H
N
ALA
V
93
R
-
C
ASN
A
455
L
H
H
ALA
V
93
R
-
C
ASN
A
455
L
H
H
SER
V
94
R
-
C
ASN
A
455
L
H
HG
SER
V
94
R
-
CB
ASN
A
455
L
H
CA
GLU
V
91
R
-
CB
ASN
A
455
L
H
C
GLU
V
91
R
-
CB
ASN
A
455
L
H
CB
GLU
V
91
R
-
CB
ASN
A
455
L
H
O
GLU
V
91
R
-
CB
ASN
A
455
L
H
HA
GLU
V
91
R
-
CB
ASN
A
455
L
H
HB2
GLU
V
91
R
-
CB
ASN
A
455
L
H
HB3
GLU
V
91
R
-
CB
ASN
A
455
L
H
N
THR
V
92
R
-
CB
ASN
A
455
L
H
HA
THR
V
92
R
-
CB
ASN
A
455
L
H
OG
SER
V
94
R
-
CB
ASN
A
455
L
H
HG
SER
V
94
R
-
CB
ASN
A
455
L
-
CE
LYS
V
95
R
-
CB
ASN
A
455
L
-
HG2
LYS
V
95
R
-
CB
ASN
A
455
L
-
HD2
LYS
V
95
R
-
CB
ASN
A
455
L
-
HE2
LYS
V
95
R
-
CB
ASN
A
455
L
-
HZ3
LYS
V
95
R
-
O
ASN
A
455
L
H
O
ASP
V
88
R
-
O
ASN
A
455
L
H
O
LEU
V
89
R
-
O
ASN
A
455
L
H
C
GLN
V
90
R
-
O
ASN
A
455
L
H
O
GLN
V
90
R
-
O
ASN
A
455
L
H
N
GLU
V
91
R
-
O
ASN
A
455
L
H
CA
GLU
V
91
R
-
O
ASN
A
455
L
H
C
GLU
V
91
R
-
O
ASN
A
455
L
H
CB
GLU
V
91
R
-
O
ASN
A
455
L
H
O
GLU
V
91
R
-
O
ASN
A
455
L
H
H
GLU
V
91
R
-
O
ASN
A
455
L
H
HA
GLU
V
91
R
-
O
ASN
A
455
L
H
HB2
GLU
V
91
R
-
O
ASN
A
455
L
H
HB3
GLU
V
91
R
-
O
ASN
A
455
L
H
N
THR
V
92
R
-
O
ASN
A
455
L
H
CA
THR
V
92
R
-
O
ASN
A
455
L
H
C
THR
V
92
R
-
O
ASN
A
455
L
H
CB
THR
V
92
R
-
O
ASN
A
455
L
H
O
THR
V
92
R
-
O
ASN
A
455
L
H
CG2
THR
V
92
R
-
O
ASN
A
455
L
H
OG1
THR
V
92
R
-
O
ASN
A
455
L
H
H
THR
V
92
R
-
O
ASN
A
455
L
H
HA
THR
V
92
R
-
O
ASN
A
455
L
H
HB
THR
V
92
R
-
O
ASN
A
455
L
H
HG1
THR
V
92
R
-
O
ASN
A
455
L
H
HG22
THR
V
92
R
-
O
ASN
A
455
L
H
HG23
THR
V
92
R
-
O
ASN
A
455
L
H
N
ALA
V
93
R
-
O
ASN
A
455
L
H
H
ALA
V
93
R
-
O
ASN
A
455
L
H
H
SER
V
94
R
-
O
ASN
A
455
L
H
HG
SER
V
94
R
-
CG
ASN
A
455
L
H
C
GLU
V
91
R
-
CG
ASN
A
455
L
H
CB
GLU
V
91
R
-
CG
ASN
A
455
L
H
O
GLU
V
91
R
-
CG
ASN
A
455
L
H
HB3
GLU
V
91
R
-
CG
ASN
A
455
L
H
OG
SER
V
94
R
-
CG
ASN
A
455
L
H
HG
SER
V
94
R
-
CG
ASN
A
455
L
-
CE
LYS
V
95
R
-
CG
ASN
A
455
L
-
HE2
LYS
V
95
R
-
CG
ASN
A
455
L
-
HZ3
LYS
V
95
R
-
ND2
ASN
A
455
L
H
HB3
GLU
V
91
R
-
ND2
ASN
A
455
L
H
OG
SER
V
94
R
-
ND2
ASN
A
455
L
H
HG
SER
V
94
R
-
ND2
ASN
A
455
L
-
CE
LYS
V
95
R
-
ND2
ASN
A
455
L
-
NZ
LYS
V
95
R
-
ND2
ASN
A
455
L
-
HE2
LYS
V
95
R
-
ND2
ASN
A
455
L
-
HE3
LYS
V
95
R
-
ND2
ASN
A
455
L
-
HZ1
LYS
V
95
R
-
ND2
ASN
A
455
L
-
HZ3
LYS
V
95
R
-
OD1
ASN
A
455
L
H
CB
GLU
V
91
R
-
OD1
ASN
A
455
L
H
CG
GLU
V
91
R
-
OD1
ASN
A
455
L
H
CD
GLU
V
91
R
-
OD1
ASN
A
455
L
H
OE1
GLU
V
91
R
-
OD1
ASN
A
455
L
H
OE2
GLU
V
91
R
-
OD1
ASN
A
455
L
H
HB2
GLU
V
91
R
-
OD1
ASN
A
455
L
H
HB3
GLU
V
91
R
-
OD1
ASN
A
455
L
H
HG2
GLU
V
91
R
-
N
THR
A
456
L
H
CA
GLN
V
90
R
-
N
THR
A
456
L
H
C
GLN
V
90
R
-
N
THR
A
456
L
H
O
GLN
V
90
R
-
N
THR
A
456
L
H
N
GLU
V
91
R
-
N
THR
A
456
L
H
CA
GLU
V
91
R
-
N
THR
A
456
L
H
C
GLU
V
91
R
-
N
THR
A
456
L
H
CB
GLU
V
91
R
-
N
THR
A
456
L
H
O
GLU
V
91
R
-
N
THR
A
456
L
H
CG
GLU
V
91
R
-
N
THR
A
456
L
H
H
GLU
V
91
R
-
N
THR
A
456
L
H
HA
GLU
V
91
R
-
N
THR
A
456
L
H
HB2
GLU
V
91
R
-
N
THR
A
456
L
H
HB3
GLU
V
91
R
-
N
THR
A
456
L
H
HG2
GLU
V
91
R
-
N
THR
A
456
L
H
HG3
GLU
V
91
R
-
N
THR
A
456
L
H
N
THR
V
92
R
-
N
THR
A
456
L
H
CA
THR
V
92
R
-
N
THR
A
456
L
H
C
THR
V
92
R
-
N
THR
A
456
L
H
H
THR
V
92
R
-
N
THR
A
456
L
H
HA
THR
V
92
R
-
N
THR
A
456
L
H
N
ALA
V
93
R
-
N
THR
A
456
L
H
H
ALA
V
93
R
-
N
THR
A
456
L
H
H
SER
V
94
R
-
N
THR
A
456
L
H
HG
SER
V
94
R
-
CA
THR
A
456
L
H
CA
GLN
V
90
R
-
CA
THR
A
456
L
H
C
GLN
V
90
R
-
CA
THR
A
456
L
H
O
GLN
V
90
R
-
CA
THR
A
456
L
H
N
GLU
V
91
R
-
CA
THR
A
456
L
H
CA
GLU
V
91
R
-
CA
THR
A
456
L
H
C
GLU
V
91
R
-
CA
THR
A
456
L
H
CB
GLU
V
91
R
-
CA
THR
A
456
L
H
O
GLU
V
91
R
-
CA
THR
A
456
L
H
H
GLU
V
91
R
-
CA
THR
A
456
L
H
HA
GLU
V
91
R
-
CA
THR
A
456
L
H
HB3
GLU
V
91
R
-
CA
THR
A
456
L
H
HG2
GLU
V
91
R
-
CA
THR
A
456
L
H
N
THR
V
92
R
-
CA
THR
A
456
L
H
C
THR
V
92
R
-
CA
THR
A
456
L
H
H
THR
V
92
R
-
CA
THR
A
456
L
H
N
ALA
V
93
R
-
CA
THR
A
456
L
H
CB
ALA
V
93
R
-
CA
THR
A
456
L
H
H
ALA
V
93
R
-
CA
THR
A
456
L
H
HB3
ALA
V
93
R
-
CA
THR
A
456
L
H
N
SER
V
94
R
-
CA
THR
A
456
L
H
H
SER
V
94
R
-
CA
THR
A
456
L
H
HB3
SER
V
94
R
-
CA
THR
A
456
L
H
HG
SER
V
94
R
-
C
THR
A
456
L
H
CA
GLN
V
90
R
-
C
THR
A
456
L
H
C
GLN
V
90
R
-
C
THR
A
456
L
H
CB
GLN
V
90
R
-
C
THR
A
456
L
H
O
GLN
V
90
R
-
C
THR
A
456
L
H
HA
GLN
V
90
R
-
C
THR
A
456
L
H
HB3
GLN
V
90
R
-
C
THR
A
456
L
H
N
GLU
V
91
R
-
C
THR
A
456
L
H
CA
GLU
V
91
R
-
C
THR
A
456
L
H
C
GLU
V
91
R
-
C
THR
A
456
L
H
O
GLU
V
91
R
-
C
THR
A
456
L
H
H
GLU
V
91
R
-
C
THR
A
456
L
H
HA
GLU
V
91
R
-
C
THR
A
456
L
H
N
THR
V
92
R
-
C
THR
A
456
L
H
C
THR
V
92
R
-
C
THR
A
456
L
H
H
THR
V
92
R
-
C
THR
A
456
L
H
N
ALA
V
93
R
-
C
THR
A
456
L
H
CA
ALA
V
93
R
-
C
THR
A
456
L
H
CB
ALA
V
93
R
-
C
THR
A
456
L
H
H
ALA
V
93
R
-
C
THR
A
456
L
H
HB1
ALA
V
93
R
-
C
THR
A
456
L
H
HB2
ALA
V
93
R
-
C
THR
A
456
L
H
HB3
ALA
V
93
R
-
C
THR
A
456
L
H
N
SER
V
94
R
-
C
THR
A
456
L
H
H
SER
V
94
R
-
CB
THR
A
456
L
H
C
GLN
V
90
R
-
CB
THR
A
456
L
H
O
GLN
V
90
R
-
CB
THR
A
456
L
H
N
GLU
V
91
R
-
CB
THR
A
456
L
H
CA
GLU
V
91
R
-
CB
THR
A
456
L
H
C
GLU
V
91
R
-
CB
THR
A
456
L
H
O
GLU
V
91
R
-
CB
THR
A
456
L
H
HA
GLU
V
91
R
-
CB
THR
A
456
L
H
N
THR
V
92
R
-
CB
THR
A
456
L
H
C
THR
V
92
R
-
CB
THR
A
456
L
H
N
ALA
V
93
R
-
CB
THR
A
456
L
H
CA
ALA
V
93
R
-
CB
THR
A
456
L
H
C
ALA
V
93
R
-
CB
THR
A
456
L
H
CB
ALA
V
93
R
-
CB
THR
A
456
L
H
H
ALA
V
93
R
-
CB
THR
A
456
L
H
HB3
ALA
V
93
R
-
CB
THR
A
456
L
H
N
SER
V
94
R
-
CB
THR
A
456
L
H
CA
SER
V
94
R
-
CB
THR
A
456
L
H
CB
SER
V
94
R
-
CB
THR
A
456
L
H
OG
SER
V
94
R
-
CB
THR
A
456
L
H
H
SER
V
94
R
-
CB
THR
A
456
L
H
HA
SER
V
94
R
-
CB
THR
A
456
L
H
HB2
SER
V
94
R
-
CB
THR
A
456
L
H
HB3
SER
V
94
R
-
CB
THR
A
456
L
H
HG
SER
V
94
R
-
O
THR
A
456
L
H
O
LEU
V
89
R
-
O
THR
A
456
L
H
CA
GLN
V
90
R
-
O
THR
A
456
L
H
C
GLN
V
90
R
-
O
THR
A
456
L
H
O
GLN
V
90
R
-
O
THR
A
456
L
H
HA
GLN
V
90
R
-
O
THR
A
456
L
H
N
GLU
V
91
R
-
O
THR
A
456
L
H
CA
GLU
V
91
R
-
O
THR
A
456
L
H
C
GLU
V
91
R
-
O
THR
A
456
L
H
O
GLU
V
91
R
-
O
THR
A
456
L
H
HA
GLU
V
91
R
-
O
THR
A
456
L
H
N
THR
V
92
R
-
O
THR
A
456
L
H
CA
THR
V
92
R
-
O
THR
A
456
L
H
C
THR
V
92
R
-
O
THR
A
456
L
H
O
THR
V
92
R
-
O
THR
A
456
L
H
H
THR
V
92
R
-
O
THR
A
456
L
H
N
ALA
V
93
R
-
O
THR
A
456
L
H
CA
ALA
V
93
R
-
O
THR
A
456
L
H
C
ALA
V
93
R
-
O
THR
A
456
L
H
CB
ALA
V
93
R
-
O
THR
A
456
L
H
H
ALA
V
93
R
-
O
THR
A
456
L
H
HA
ALA
V
93
R
-
O
THR
A
456
L
H
HB1
ALA
V
93
R
-
O
THR
A
456
L
H
HB2
ALA
V
93
R
-
O
THR
A
456
L
H
HB3
ALA
V
93
R
-
O
THR
A
456
L
H
N
SER
V
94
R
-
O
THR
A
456
L
H
H
SER
V
94
R
-
CG2
THR
A
456
L
H
CA
GLU
V
91
R
-
CG2
THR
A
456
L
H
C
GLU
V
91
R
-
CG2
THR
A
456
L
H
O
GLU
V
91
R
-
CG2
THR
A
456
L
H
HA
GLU
V
91
R
-
CG2
THR
A
456
L
H
HG2
GLU
V
91
R
-
CG2
THR
A
456
L
H
N
SER
V
94
R
-
CG2
THR
A
456
L
H
CA
SER
V
94
R
-
CG2
THR
A
456
L
H
CB
SER
V
94
R
-
CG2
THR
A
456
L
H
OG
SER
V
94
R
-
CG2
THR
A
456
L
H
H
SER
V
94
R
-
CG2
THR
A
456
L
H
HB2
SER
V
94
R
-
CG2
THR
A
456
L
H
HB3
SER
V
94
R
-
CG2
THR
A
456
L
H
HG
SER
V
94
R
-
OG1
THR
A
456
L
H
O
GLN
V
90
R
-
OG1
THR
A
456
L
H
CA
GLU
V
91
R
-
OG1
THR
A
456
L
H
C
GLU
V
91
R
-
OG1
THR
A
456
L
H
O
GLU
V
91
R
-
OG1
THR
A
456
L
H
HA
GLU
V
91
R
-
OG1
THR
A
456
L
H
N
THR
V
92
R
-
OG1
THR
A
456
L
H
CA
THR
V
92
R
-
OG1
THR
A
456
L
H
C
THR
V
92
R
-
OG1
THR
A
456
L
H
O
THR
V
92
R
-
OG1
THR
A
456
L
H
HA
THR
V
92
R
-
OG1
THR
A
456
L
H
N
ALA
V
93
R
-
OG1
THR
A
456
L
H
CA
ALA
V
93
R
-
OG1
THR
A
456
L
H
C
ALA
V
93
R
-
OG1
THR
A
456
L
H
H
ALA
V
93
R
-
OG1
THR
A
456
L
H
HB3
ALA
V
93
R
-
OG1
THR
A
456
L
H
N
SER
V
94
R
-
OG1
THR
A
456
L
H
CA
SER
V
94
R
-
OG1
THR
A
456
L
H
C
SER
V
94
R
-
OG1
THR
A
456
L
H
CB
SER
V
94
R
-
OG1
THR
A
456
L
H
OG
SER
V
94
R
-
OG1
THR
A
456
L
H
H
SER
V
94
R
-
OG1
THR
A
456
L
H
HA
SER
V
94
R
-
OG1
THR
A
456
L
H
HB2
SER
V
94
R
-
OG1
THR
A
456
L
H
HB3
SER
V
94
R
-
OG1
THR
A
456
L
H
HG
SER
V
94
R
-
OG1
THR
A
456
L
-
N
LYS
V
95
R
-
OG1
THR
A
456
L
-
H
LYS
V
95
R
-
OG1
THR
A
456
L
-
HG2
LYS
V
95
R
-
N
GLN
A
457
L
H
CA
GLN
V
90
R
-
N
GLN
A
457
L
H
C
GLN
V
90
R
-
N
GLN
A
457
L
H
CB
GLN
V
90
R
-
N
GLN
A
457
L
H
O
GLN
V
90
R
-
N
GLN
A
457
L
H
HA
GLN
V
90
R
-
N
GLN
A
457
L
H
HB2
GLN
V
90
R
-
N
GLN
A
457
L
H
HB3
GLN
V
90
R
-
N
GLN
A
457
L
H
HG2
GLN
V
90
R
-
N
GLN
A
457
L
H
N
GLU
V
91
R
-
N
GLN
A
457
L
H
CA
GLU
V
91
R
-
N
GLN
A
457
L
H
H
GLU
V
91
R
-
N
GLN
A
457
L
H
HA
GLU
V
91
R
-
N
GLN
A
457
L
H
N
ALA
V
93
R
-
N
GLN
A
457
L
H
CB
ALA
V
93
R
-
N
GLN
A
457
L
H
H
ALA
V
93
R
-
N
GLN
A
457
L
H
HB2
ALA
V
93
R
-
N
GLN
A
457
L
H
HB3
ALA
V
93
R
-
CA
GLN
A
457
L
H
CA
GLN
V
90
R
-
CA
GLN
A
457
L
H
C
GLN
V
90
R
-
CA
GLN
A
457
L
H
CB
GLN
V
90
R
-
CA
GLN
A
457
L
H
O
GLN
V
90
R
-
CA
GLN
A
457
L
H
HA
GLN
V
90
R
-
CA
GLN
A
457
L
H
HB3
GLN
V
90
R
-
CA
GLN
A
457
L
H
HG2
GLN
V
90
R
-
CA
GLN
A
457
L
H
CB
ALA
V
93
R
-
CA
GLN
A
457
L
H
HB1
ALA
V
93
R
-
CA
GLN
A
457
L
H
HB2
ALA
V
93
R
-
CA
GLN
A
457
L
H
HB3
ALA
V
93
R
-
C
GLN
A
457
L
H
O
GLN
V
90
R
-
C
GLN
A
457
L
H
HA
GLN
V
90
R
-
C
GLN
A
457
L
H
HB3
GLN
V
90
R
-
C
GLN
A
457
L
H
HG2
GLN
V
90
R
-
C
GLN
A
457
L
H
CB
ALA
V
93
R
-
C
GLN
A
457
L
H
HB1
ALA
V
93
R
-
C
GLN
A
457
L
H
HB2
ALA
V
93
R
-
C
GLN
A
457
L
H
HB3
ALA
V
93
R
-
CB
GLN
A
457
L
H
O
GLN
V
90
R
-
CB
GLN
A
457
L
H
CB
ALA
V
93
R
-
CB
GLN
A
457
L
H
HB1
ALA
V
93
R
-
CB
GLN
A
457
L
H
HB2
ALA
V
93
R
-
CB
GLN
A
457
L
H
HB3
ALA
V
93
R
-
O
GLN
A
457
L
H
HG22
ILE
V
39
R
-
O
GLN
A
457
L
H
HD12
ILE
V
39
R
-
O
GLN
A
457
L
H
O
GLN
V
90
R
-
O
GLN
A
457
L
H
HA
GLN
V
90
R
-
O
GLN
A
457
L
H
HG2
GLN
V
90
R
-
O
GLN
A
457
L
H
CB
ALA
V
93
R
-
O
GLN
A
457
L
H
HB1
ALA
V
93
R
-
O
GLN
A
457
L
H
HB2
ALA
V
93
R
-
O
GLN
A
457
L
H
HB3
ALA
V
93
R
-
CG
GLN
A
457
L
H
HB3
ALA
V
93
R
-
N
LEU
A
458
L
H
HB3
GLN
V
90
R
-
N
LEU
A
458
L
H
HG2
GLN
V
90
R
-
C
LEU
A
458
L
H
HG22
ILE
V
39
R
-
C
LEU
A
458
L
H
OD1
ASP
V
40
R
-
O
LEU
A
458
L
H
CG2
ILE
V
39
R
-
O
LEU
A
458
L
H
HB
ILE
V
39
R
-
O
LEU
A
458
L
H
HG21
ILE
V
39
R
-
O
LEU
A
458
L
H
HG22
ILE
V
39
R
-
O
LEU
A
458
L
H
OD1
ASP
V
40
R
-
N
GLY
A
459
L
H
OD1
ASP
V
40
R
-
CA
GLY
A
459
L
H
HH21
ARG
V
36
R
-
CA
GLY
A
459
L
H
CG
ASP
V
40
R
-
CA
GLY
A
459
L
H
OD1
ASP
V
40
R
-
CA
GLY
A
459
L
H
OD2
ASP
V
40
R
-
C
GLY
A
459
L
H
CA
ASP
V
40
R
-
C
GLY
A
459
L
H
CB
ASP
V
40
R
-
C
GLY
A
459
L
H
CG
ASP
V
40
R
-
C
GLY
A
459
L
H
OD1
ASP
V
40
R
-
C
GLY
A
459
L
H
OD2
ASP
V
40
R
-
C
GLY
A
459
L
H
HA
ASP
V
40
R
-
C
GLY
A
459
L
H
HB3
ASP
V
40
R
-
C
GLY
A
459
L
H
HG23
VAL
V
43
R
-
O
GLY
A
459
L
H
C
ILE
V
39
R
-
O
GLY
A
459
L
H
HG21
ILE
V
39
R
-
O
GLY
A
459
L
H
N
ASP
V
40
R
-
O
GLY
A
459
L
H
CA
ASP
V
40
R
-
O
GLY
A
459
L
H
C
ASP
V
40
R
-
O
GLY
A
459
L
H
CB
ASP
V
40
R
-
O
GLY
A
459
L
H
CG
ASP
V
40
R
-
O
GLY
A
459
L
H
OD1
ASP
V
40
R
-
O
GLY
A
459
L
H
OD2
ASP
V
40
R
-
O
GLY
A
459
L
H
H
ASP
V
40
R
-
O
GLY
A
459
L
H
HA
ASP
V
40
R
-
O
GLY
A
459
L
H
HB3
ASP
V
40
R
-
O
GLY
A
459
L
H
CG2
VAL
V
43
R
-
O
GLY
A
459
L
H
HB
VAL
V
43
R
-
O
GLY
A
459
L
H
HG21
VAL
V
43
R
-
O
GLY
A
459
L
H
HG23
VAL
V
43
R
-
N
GLY
A
460
L
H
CG
ASP
V
40
R
-
N
GLY
A
460
L
H
OD1
ASP
V
40
R
-
N
GLY
A
460
L
H
HA
ASP
V
40
R
-
CA
GLY
A
460
L
H
CG
ASP
V
40
R
-
CA
GLY
A
460
L
H
OD1
ASP
V
40
R
-
CA
GLY
A
460
L
H
HA
ASP
V
40
R
-
CA
GLY
A
460
L
H
HB3
ASP
V
40
R
-
CA
GLY
A
460
L
H
HB
VAL
V
43
R
-
CA
GLY
A
460
L
H
HG12
VAL
V
43
R
-
C
GLY
A
460
L
H
CA
ASP
V
40
R
-
C
GLY
A
460
L
H
CB
ASP
V
40
R
-
C
GLY
A
460
L
H
O
ASP
V
40
R
-
C
GLY
A
460
L
H
CG
ASP
V
40
R
-
C
GLY
A
460
L
H
OD1
ASP
V
40
R
-
C
GLY
A
460
L
H
HA
ASP
V
40
R
-
C
GLY
A
460
L
H
HB3
ASP
V
40
R
-
O
GLY
A
460
L
H
CB
ASP
V
40
R
-
O
GLY
A
460
L
H
CG
ASP
V
40
R
-
O
GLY
A
460
L
H
OD1
ASP
V
40
R
-
O
GLY
A
460
L
H
OD2
ASP
V
40
R
-
O
GLY
A
460
L
H
HB3
ASP
V
40
R
-
N
GLY
A
461
L
H
CA
ASP
V
40
R
-
N
GLY
A
461
L
H
C
ASP
V
40
R
-
N
GLY
A
461
L
H
CB
ASP
V
40
R
-
N
GLY
A
461
L
H
O
ASP
V
40
R
-
N
GLY
A
461
L
H
HA
ASP
V
40
R
-
N
GLY
A
461
L
H
HB3
ASP
V
40
R
-
N
GLY
A
461
L
H
OG
SER
V
44
R
-
N
GLY
A
461
L
H
HG
SER
V
44
R
-
CA
GLY
A
461
L
H
CB
ASP
V
40
R
-
CA
GLY
A
461
L
H
O
ASP
V
40
R
-
CA
GLY
A
461
L
H
HB3
ASP
V
40
R
-
CA
GLY
A
461
L
H
OG
SER
V
44
R
-
CA
GLY
A
461
L
H
HG
SER
V
44
R
-
C
GLY
A
461
L
H
O
ASP
V
40
R
-
C
GLY
A
461
L
H
HB3
ASP
V
40
R
-
C
GLY
A
461
L
H
HA
THR
V
41
R
-
C
GLY
A
461
L
H
CB
SER
V
44
R
-
C
GLY
A
461
L
H
OG
SER
V
44
R
-
C
GLY
A
461
L
H
HB2
SER
V
44
R
-
C
GLY
A
461
L
H
HG
SER
V
44
R
-
O
GLY
A
461
L
H
C
ASP
V
40
R
-
O
GLY
A
461
L
H
O
ASP
V
40
R
-
O
GLY
A
461
L
H
HB3
ASP
V
40
R
-
O
GLY
A
461
L
H
N
THR
V
41
R
-
O
GLY
A
461
L
H
CA
THR
V
41
R
-
O
GLY
A
461
L
H
HA
THR
V
41
R
-
O
GLY
A
461
L
H
HG23
THR
V
41
R
-
O
GLY
A
461
L
H
CB
SER
V
44
R
-
O
GLY
A
461
L
H
OG
SER
V
44
R
-
O
GLY
A
461
L
H
HB2
SER
V
44
R
-
O
GLY
A
461
L
H
HB3
SER
V
44
R
-
O
GLY
A
461
L
H
HG
SER
V
44
R
-
N
THR
A
462
L
H
CB
SER
V
44
R
-
N
THR
A
462
L
H
OG
SER
V
44
R
-
N
THR
A
462
L
H
HB3
SER
V
44
R
-
N
THR
A
462
L
H
HG
SER
V
44
R
-
CA
THR
A
462
L
H
CB
SER
V
44
R
-
CA
THR
A
462
L
H
OG
SER
V
44
R
-
CA
THR
A
462
L
H
HB2
SER
V
44
R
-
CA
THR
A
462
L
H
HB3
SER
V
44
R
-
CA
THR
A
462
L
H
HG
SER
V
44
R
-
CB
THR
A
462
L
H
OG
SER
V
44
R
-
CB
THR
A
462
L
H
HB3
SER
V
44
R
-
CG2
THR
A
462
L
H
CB
SER
V
44
R
-
CG2
THR
A
462
L
H
OG
SER
V
44
R
-
CG2
THR
A
462
L
H
HB3
SER
V
44
R
-
O
GLY
A
463
L
H
HH12
ARG
V
174
R
-
CA
ALA
A
464
L
H
HH12
ARG
V
174
R
-
CB
ALA
A
464
L
H
HZ
PHE
V
163
R
-
N
PRO
A
465
L
H
HE2
PHE
V
163
R
-
CA
PRO
A
465
L
H
HE2
PHE
V
163
R
-
CA
PRO
A
465
L
H
HH22
ARG
V
174
R
-
C
PRO
A
465
L
H
HE22
GLN
V
170
R
-
C
PRO
A
465
L
H
NH2
ARG
V
174
R
-
C
PRO
A
465
L
H
HH21
ARG
V
174
R
-
C
PRO
A
465
L
H
HH22
ARG
V
174
R
-
CB
PRO
A
465
L
H
CD2
PHE
V
163
R
-
CB
PRO
A
465
L
H
CE2
PHE
V
163
R
-
CB
PRO
A
465
L
H
CZ
PHE
V
163
R
-
CB
PRO
A
465
L
H
HD2
PHE
V
163
R
-
CB
PRO
A
465
L
H
HE2
PHE
V
163
R
-
CB
PRO
A
465
L
H
HZ
PHE
V
163
R
-
CB
PRO
A
465
L
H
HE22
GLN
V
170
R
-
O
PRO
A
465
L
H
HH22
ARG
V
174
R
-
CG
PRO
A
465
L
H
CD2
PHE
V
163
R
-
CG
PRO
A
465
L
H
CE2
PHE
V
163
R
-
CG
PRO
A
465
L
H
HD2
PHE
V
163
R
-
CG
PRO
A
465
L
H
HE2
PHE
V
163
R
-
CD
PRO
A
465
L
H
HE2
PHE
V
163
R
-
N
HIS
A
466
L
H
CD
GLN
V
170
R
-
N
HIS
A
466
L
H
NE2
GLN
V
170
R
-
N
HIS
A
466
L
H
OE1
GLN
V
170
R
-
N
HIS
A
466
L
H
HE21
GLN
V
170
R
-
N
HIS
A
466
L
H
HE22
GLN
V
170
R
-
N
HIS
A
466
L
H
NH2
ARG
V
174
R
-
N
HIS
A
466
L
H
HH21
ARG
V
174
R
-
N
HIS
A
466
L
H
HH22
ARG
V
174
R
-
CA
HIS
A
466
L
H
CD
GLN
V
170
R
-
CA
HIS
A
466
L
H
NE2
GLN
V
170
R
-
CA
HIS
A
466
L
H
HE21
GLN
V
170
R
-
CA
HIS
A
466
L
H
HE22
GLN
V
170
R
-
CA
HIS
A
466
L
H
HH21
ARG
V
174
R
-
C
HIS
A
466
L
H
CD
GLN
V
170
R
-
C
HIS
A
466
L
H
NE2
GLN
V
170
R
-
C
HIS
A
466
L
H
OE1
GLN
V
170
R
-
C
HIS
A
466
L
H
HG3
GLN
V
170
R
-
C
HIS
A
466
L
H
HE21
GLN
V
170
R
-
C
HIS
A
466
L
H
HE22
GLN
V
170
R
-
CB
HIS
A
466
L
H
HH21
ARG
V
174
R
-
O
HIS
A
466
L
-
HG13
VAL
V
166
R
-
O
HIS
A
466
L
H
CG
GLN
V
170
R
-
O
HIS
A
466
L
H
CD
GLN
V
170
R
-
O
HIS
A
466
L
H
NE2
GLN
V
170
R
-
O
HIS
A
466
L
H
OE1
GLN
V
170
R
-
O
HIS
A
466
L
H
HG2
GLN
V
170
R
-
O
HIS
A
466
L
H
HG3
GLN
V
170
R
-
O
HIS
A
466
L
H
HE21
GLN
V
170
R
-
O
HIS
A
466
L
H
HE22
GLN
V
170
R
-
N
SER
A
467
L
H
NE2
GLN
V
170
R
-
N
SER
A
467
L
H
HE21
GLN
V
170
R
-
N
SER
A
467
L
H
HE22
GLN
V
170
R
-
CA
SER
A
467
L
-
HG23
VAL
V
166
R
-
CA
SER
A
467
L
H
NE2
GLN
V
170
R
-
CA
SER
A
467
L
H
HE21
GLN
V
170
R
-
CA
SER
A
467
L
H
HE22
GLN
V
170
R
-
C
SER
A
467
L
-
HG23
VAL
V
166
R
-
C
SER
A
467
L
H
HE21
GLN
V
170
R
-
CB
SER
A
467
L
H
HE21
GLN
V
170
R
-
CB
SER
A
467
L
H
HE22
GLN
V
170
R
-
OG
SER
A
467
L
H
HE22
GLN
V
170
R
-
N
ALA
A
468
L
-
HG23
VAL
V
166
R
-
N
ALA
A
468
L
H
HE21
GLN
V
170
R
-
C
ALA
A
468
L
H
HE1
TYR
V
125
R
-
C
ALA
A
468
L
-
HG23
VAL
V
166
R
-
O
ALA
A
468
L
H
CE1
TYR
V
125
R
-
O
ALA
A
468
L
H
OH
TYR
V
125
R
-
O
ALA
A
468
L
H
CZ
TYR
V
125
R
-
O
ALA
A
468
L
H
HE1
TYR
V
125
R
-
O
ALA
A
468
L
H
HH
TYR
V
125
R
-
O
ALA
A
468
L
-
CG2
VAL
V
166
R
-
O
ALA
A
468
L
-
HG21
VAL
V
166
R
-
O
ALA
A
468
L
-
HG23
VAL
V
166
R
-
O
ALA
A
468
L
H
HD22
LEU
V
169
R
-
N
SER
A
469
L
H
HE1
TYR
V
125
R
-
N
SER
A
469
L
H
HH
TYR
V
125
R
-
C
SER
A
469
L
H
HE1
TYR
V
125
R
-
C
SER
A
469
L
H
CD1
LEU
V
129
R
-
C
SER
A
469
L
H
HD11
LEU
V
129
R
-
C
SER
A
469
L
H
HD12
LEU
V
129
R
-
C
SER
A
469
L
H
HD13
LEU
V
129
R
-
O
SER
A
469
L
H
CE1
TYR
V
125
R
-
O
SER
A
469
L
H
HE1
TYR
V
125
R
-
O
SER
A
469
L
H
HD11
LEU
V
129
R
-
O
SER
A
469
L
H
HD13
LEU
V
129
R
-
N
SER
A
470
L
H
CD1
LEU
V
129
R
-
N
SER
A
470
L
H
HD11
LEU
V
129
R
-
N
SER
A
470
L
H
HD12
LEU
V
129
R
-
N
SER
A
470
L
H
HD13
LEU
V
129
R
-
CA
SER
A
470
L
H
HD12
LEU
V
129
R
-
C
SER
A
470
L
H
HD12
LEU
V
129
R
-
O
SER
A
470
L
H
HB3
LEU
V
129
R
-
O
SER
A
470
L
H
HD12
LEU
V
129
R
-
O
SER
A
470
L
H
HD13
LEU
V
129
R
-
C
ALA
A
471
L
H
HD21
LEU
V
130
R
-
C
ALA
A
471
L
H
HD23
LEU
V
130
R
-
C
ALA
A
471
L
H
OD2
ASP
V
133
R
-
O
ALA
A
471
L
H
HB3
LEU
V
129
R
-
O
ALA
A
471
L
H
CG
LEU
V
130
R
-
O
ALA
A
471
L
H
CD2
LEU
V
130
R
-
O
ALA
A
471
L
H
HA
LEU
V
130
R
-
O
ALA
A
471
L
H
HG
LEU
V
130
R
-
O
ALA
A
471
L
H
HD21
LEU
V
130
R
-
O
ALA
A
471
L
H
HD23
LEU
V
130
R
-
O
ALA
A
471
L
H
OD2
ASP
V
133
R
-
N
SER
A
472
L
H
HD21
LEU
V
130
R
-
N
SER
A
472
L
H
CG
ASP
V
133
R
-
N
SER
A
472
L
H
OD2
ASP
V
133
R
-
N
SER
A
472
L
H
HB2
ASP
V
133
R
-
N
SER
A
472
L
H
HB3
ASP
V
133
R
-
CA
SER
A
472
L
H
CG
ASP
V
133
R
-
CA
SER
A
472
L
H
OD2
ASP
V
133
R
-
CA
SER
A
472
L
H
HB2
ASP
V
133
R
-
C
SER
A
472
L
H
HD21
LEU
V
130
R
-
CB
SER
A
472
L
H
OD2
ASP
V
133
R
-
O
SER
A
472
L
H
CD2
LEU
V
130
R
-
O
SER
A
472
L
H
HD21
LEU
V
130
R
-
O
SER
A
472
L
H
HD22
LEU
V
130
R
-
O
SER
A
472
L
H
HD23
LEU
V
130
R
-
N
LEU
A
473
L
H
NZ
LYS
V
137
R
-
N
LEU
A
473
L
H
HZ1
LYS
V
137
R
-
CA
LEU
A
473
L
H
CE
LYS
V
137
R
-
CA
LEU
A
473
L
H
NZ
LYS
V
137
R
-
CA
LEU
A
473
L
H
HE2
LYS
V
137
R
-
CA
LEU
A
473
L
H
HE3
LYS
V
137
R
-
CA
LEU
A
473
L
H
HZ1
LYS
V
137
R
-
CA
LEU
A
473
L
H
HZ2
LYS
V
137
R
-
C
LEU
A
473
L
H
CD
LYS
V
137
R
-
C
LEU
A
473
L
H
CE
LYS
V
137
R
-
C
LEU
A
473
L
H
NZ
LYS
V
137
R
-
C
LEU
A
473
L
H
HE2
LYS
V
137
R
-
C
LEU
A
473
L
H
HE3
LYS
V
137
R
-
C
LEU
A
473
L
H
HZ1
LYS
V
137
R
-
C
LEU
A
473
L
H
HZ2
LYS
V
137
R
-
C
LEU
A
473
L
H
HZ3
LYS
V
137
R
-
CB
LEU
A
473
L
H
HZ1
LYS
V
137
R
-
O
LEU
A
473
L
H
CD
LYS
V
137
R
-
O
LEU
A
473
L
H
CE
LYS
V
137
R
-
O
LEU
A
473
L
H
NZ
LYS
V
137
R
-
O
LEU
A
473
L
H
HD3
LYS
V
137
R
-
O
LEU
A
473
L
H
HE2
LYS
V
137
R
-
O
LEU
A
473
L
H
HE3
LYS
V
137
R
-
O
LEU
A
473
L
H
HZ1
LYS
V
137
R
-
O
LEU
A
473
L
H
HZ2
LYS
V
137
R
-
O
LEU
A
473
L
H
HZ3
LYS
V
137
R
-
N
HIS
A
474
L
H
CG
LYS
V
137
R
-
N
HIS
A
474
L
H
CD
LYS
V
137
R
-
N
HIS
A
474
L
H
CE
LYS
V
137
R
-
N
HIS
A
474
L
H
NZ
LYS
V
137
R
-
N
HIS
A
474
L
H
HG2
LYS
V
137
R
-
N
HIS
A
474
L
H
HG3
LYS
V
137
R
-
N
HIS
A
474
L
H
HD2
LYS
V
137
R
-
N
HIS
A
474
L
H
HD3
LYS
V
137
R
-
N
HIS
A
474
L
H
HE2
LYS
V
137
R
-
N
HIS
A
474
L
H
HE3
LYS
V
137
R
-
N
HIS
A
474
L
H
HZ1
LYS
V
137
R
-
N
HIS
A
474
L
H
HZ2
LYS
V
137
R
-
N
HIS
A
474
L
H
HZ3
LYS
V
137
R
-
CA
HIS
A
474
L
H
CG
LYS
V
137
R
-
CA
HIS
A
474
L
H
CD
LYS
V
137
R
-
CA
HIS
A
474
L
H
CE
LYS
V
137
R
-
CA
HIS
A
474
L
H
NZ
LYS
V
137
R
-
CA
HIS
A
474
L
H
HG2
LYS
V
137
R
-
CA
HIS
A
474
L
H
HG3
LYS
V
137
R
-
CA
HIS
A
474
L
H
HD2
LYS
V
137
R
-
CA
HIS
A
474
L
H
HD3
LYS
V
137
R
-
CA
HIS
A
474
L
H
HE2
LYS
V
137
R
-
CA
HIS
A
474
L
H
HE3
LYS
V
137
R
-
CA
HIS
A
474
L
H
HZ1
LYS
V
137
R
-
CA
HIS
A
474
L
H
HZ2
LYS
V
137
R
-
CA
HIS
A
474
L
H
HZ3
LYS
V
137
R
-
C
HIS
A
474
L
H
CG
LYS
V
137
R
-
C
HIS
A
474
L
H
CD
LYS
V
137
R
-
C
HIS
A
474
L
H
CE
LYS
V
137
R
-
C
HIS
A
474
L
H
NZ
LYS
V
137
R
-
C
HIS
A
474
L
H
HB3
LYS
V
137
R
-
C
HIS
A
474
L
H
HG2
LYS
V
137
R
-
C
HIS
A
474
L
H
HD2
LYS
V
137
R
-
C
HIS
A
474
L
H
HD3
LYS
V
137
R
-
C
HIS
A
474
L
H
HE2
LYS
V
137
R
-
C
HIS
A
474
L
H
HE3
LYS
V
137
R
-
C
HIS
A
474
L
H
HZ1
LYS
V
137
R
-
C
HIS
A
474
L
H
HZ2
LYS
V
137
R
-
C
HIS
A
474
L
H
HZ3
LYS
V
137
R
-
CB
HIS
A
474
L
H
CE
LYS
V
137
R
-
CB
HIS
A
474
L
H
HG2
LYS
V
137
R
-
CB
HIS
A
474
L
H
HE2
LYS
V
137
R
-
O
HIS
A
474
L
H
CD
LYS
V
137
R
-
O
HIS
A
474
L
H
CE
LYS
V
137
R
-
O
HIS
A
474
L
H
NZ
LYS
V
137
R
-
O
HIS
A
474
L
H
HD3
LYS
V
137
R
-
O
HIS
A
474
L
H
HE2
LYS
V
137
R
-
O
HIS
A
474
L
H
HZ1
LYS
V
137
R
-
O
HIS
A
474
L
H
HZ2
LYS
V
137
R
-
CG
HIS
A
474
L
H
HE2
LYS
V
137
R
-
ND1
HIS
A
474
L
H
HE2
LYS
V
137
R
-
N
SER
A
475
L
H
CB
LYS
V
137
R
-
N
SER
A
475
L
H
CG
LYS
V
137
R
-
N
SER
A
475
L
H
CD
LYS
V
137
R
-
N
SER
A
475
L
H
CE
LYS
V
137
R
-
N
SER
A
475
L
H
NZ
LYS
V
137
R
-
N
SER
A
475
L
H
HB2
LYS
V
137
R
-
N
SER
A
475
L
H
HB3
LYS
V
137
R
-
N
SER
A
475
L
H
HG2
LYS
V
137
R
-
N
SER
A
475
L
H
HG3
LYS
V
137
R
-
N
SER
A
475
L
H
HD2
LYS
V
137
R
-
N
SER
A
475
L
H
HD3
LYS
V
137
R
-
N
SER
A
475
L
H
HE2
LYS
V
137
R
-
N
SER
A
475
L
H
HE3
LYS
V
137
R
-
N
SER
A
475
L
H
HZ2
LYS
V
137
R
-
CA
SER
A
475
L
H
CB
LYS
V
137
R
-
CA
SER
A
475
L
H
CG
LYS
V
137
R
-
CA
SER
A
475
L
H
CD
LYS
V
137
R
-
CA
SER
A
475
L
H
HB3
LYS
V
137
R
-
CA
SER
A
475
L
H
HG2
LYS
V
137
R
-
CA
SER
A
475
L
H
HD3
LYS
V
137
R
-
C
SER
A
475
L
H
CA
LYS
V
137
R
-
C
SER
A
475
L
H
CB
LYS
V
137
R
-
C
SER
A
475
L
H
O
LYS
V
137
R
-
C
SER
A
475
L
H
CG
LYS
V
137
R
-
C
SER
A
475
L
H
HA
LYS
V
137
R
-
C
SER
A
475
L
H
HB3
LYS
V
137
R
-
C
SER
A
475
L
H
HG2
LYS
V
137
R
-
C
SER
A
475
L
H
HD3
LYS
V
137
R
-
CB
SER
A
475
L
H
HB3
LYS
V
137
R
-
CB
SER
A
475
L
H
HD3
LYS
V
137
R
-
O
SER
A
475
L
H
CA
LYS
V
137
R
-
O
SER
A
475
L
H
C
LYS
V
137
R
-
O
SER
A
475
L
H
CB
LYS
V
137
R
-
O
SER
A
475
L
H
O
LYS
V
137
R
-
O
SER
A
475
L
H
CG
LYS
V
137
R
-
O
SER
A
475
L
H
HA
LYS
V
137
R
-
O
SER
A
475
L
H
HB2
LYS
V
137
R
-
O
SER
A
475
L
H
HB3
LYS
V
137
R
-
O
SER
A
475
L
H
HG2
LYS
V
137
R
-
O
SER
A
475
L
H
HD3
LYS
V
137
R
-
N
GLU
A
476
L
H
CA
LYS
V
137
R
-
N
GLU
A
476
L
H
CB
LYS
V
137
R
-
N
GLU
A
476
L
H
O
LYS
V
137
R
-
N
GLU
A
476
L
H
HA
LYS
V
137
R
-
N
GLU
A
476
L
H
HB3
LYS
V
137
R
-
N
GLU
A
476
L
H
HG2
LYS
V
137
R
-
N
ARG
A
477
L
H
HG2
PRO
V
139
R
-
N
ARG
A
477
L
H
HD2
PRO
V
139
R
-
CA
ARG
A
477
L
H
CG
PRO
V
139
R
-
CA
ARG
A
477
L
H
HG2
PRO
V
139
R
-
CA
ARG
A
477
L
H
HG3
PRO
V
139
R
-
C
ARG
A
477
L
H
CG
PRO
V
139
R
-
C
ARG
A
477
L
H
CD
PRO
V
139
R
-
C
ARG
A
477
L
H
HG2
PRO
V
139
R
-
C
ARG
A
477
L
H
HG3
PRO
V
139
R
-
C
ARG
A
477
L
H
HD2
PRO
V
139
R
-
C
ARG
A
477
L
H
HD3
PRO
V
139
R
-
O
ARG
A
477
L
H
CB
PRO
V
139
R
-
O
ARG
A
477
L
H
CG
PRO
V
139
R
-
O
ARG
A
477
L
H
CD
PRO
V
139
R
-
O
ARG
A
477
L
H
HB2
PRO
V
139
R
-
O
ARG
A
477
L
H
HG2
PRO
V
139
R
-
O
ARG
A
477
L
H
HG3
PRO
V
139
R
-
O
ARG
A
477
L
H
HD2
PRO
V
139
R
-
O
ARG
A
477
L
H
HD3
PRO
V
139
R
-
N
PRO
A
478
L
H
CB
PRO
V
139
R
-
N
PRO
A
478
L
H
CG
PRO
V
139
R
-
N
PRO
A
478
L
H
HB2
PRO
V
139
R
-
N
PRO
A
478
L
H
HB3
PRO
V
139
R
-
N
PRO
A
478
L
H
HG2
PRO
V
139
R
-
N
PRO
A
478
L
H
HG3
PRO
V
139
R
-
CA
PRO
A
478
L
H
CB
PRO
V
139
R
-
CA
PRO
A
478
L
H
CG
PRO
V
139
R
-
CA
PRO
A
478
L
H
HB2
PRO
V
139
R
-
CA
PRO
A
478
L
H
HB3
PRO
V
139
R
-
CA
PRO
A
478
L
H
HG2
PRO
V
139
R
-
CA
PRO
A
478
L
H
HG3
PRO
V
139
R
-
C
PRO
A
478
L
H
HB2
PRO
V
139
R
-
C
PRO
A
478
L
H
HG2
PRO
V
139
R
-
CB
PRO
A
478
L
H
CB
PRO
V
139
R
-
CB
PRO
A
478
L
H
CG
PRO
V
139
R
-
CB
PRO
A
478
L
H
HB2
PRO
V
139
R
-
CB
PRO
A
478
L
H
HB3
PRO
V
139
R
-
CB
PRO
A
478
L
H
HG2
PRO
V
139
R
-
CB
PRO
A
478
L
H
HG3
PRO
V
139
R
-
CG
PRO
A
478
L
H
CG
PRO
V
139
R
-
CG
PRO
A
478
L
H
HB3
PRO
V
139
R
-
CG
PRO
A
478
L
H
HG2
PRO
V
139
R
-
CG
PRO
A
478
L
H
HG3
PRO
V
139
R
-
CD
PRO
A
478
L
H
HG2
PRO
V
139
R
-
CD
PRO
A
478
L
H
HG3
PRO
V
139
R
-
N
LEU
A
479
L
H
HB2
PRO
V
139
R
-
N
LEU
A
479
L
H
NH2
ARG
V
143
R
-
N
LEU
A
479
L
H
HH21
ARG
V
143
R
-
N
LEU
A
479
L
H
HH22
ARG
V
143
R
-
CA
LEU
A
479
L
H
HH21
ARG
V
143
R
-
CA
LEU
A
479
L
H
HH22
ARG
V
143
R
-
C
LEU
A
479
L
H
NH2
ARG
V
143
R
-
C
LEU
A
479
L
H
HH22
ARG
V
143
R
-
O
LEU
A
479
L
H
HH22
ARG
V
143
R
-
N
SER
A
480
L
H
NH2
ARG
V
143
R
-
N
SER
A
480
L
H
HH22
ARG
V
143
R
H
CD2
LEU
A
620
L
H
HH21
ARG
V
429
R
H
CD
GLN
A
623
L
H
HE22
GLN
V
419
R
H
NE2
GLN
A
623
L
H
HE22
GLN
V
419
R
H
OE1
GLN
A
623
L
H
NE2
GLN
V
419
R
H
OE1
GLN
A
623
L
H
HE21
GLN
V
419
R
H
OE1
GLN
A
623
L
H
HE22
GLN
V
419
R
H
ND2
ASN
A
624
L
H
HD22
LEU
V
342
R
-
NH2
ARG
A
636
L
-
HB3
PRO
V
64
R
-
N
GLY
A
637
L
H
HG3
GLN
V
65
R
-
CA
GLY
A
637
L
H
HG3
GLN
V
65
R
-
CE2
PHE
A
640
L
H
NE2
GLN
V
65
R
-
CE2
PHE
A
640
L
H
HE21
GLN
V
65
R
-
CE2
PHE
A
640
L
H
HE22
GLN
V
65