A1IGU5_Q99961.1
contacts (residues)
Receptor
R: Secondary Structure
R: Residue
R; Chain
R: Residue number
Ligand
L: Secondary Structure
L: Residue
L: Chain
L: Residue Number
R
H
TRP
A
278
L
-
GLU
V
329
R
H
LEU
A
281
L
-
TYR
V
315
R
H
CYS
A
282
L
-
TYR
V
315
R
H
GLU
A
285
L
-
TYR
V
315
R
H
GLU
A
285
L
H
TYR
V
359
R
H
ASN
A
289
L
H
TYR
V
359
R
-
LEU
A
310
L
H
ASN
V
109
R
-
LEU
A
310
L
H
PHE
V
110
R
-
LEU
A
310
L
H
MET
V
244
R
-
LEU
A
310
L
H
ALA
V
247
R
-
ASP
A
311
L
H
MET
V
244
R
-
ASP
A
311
L
H
ALA
V
247
R
-
ASP
A
311
L
H
SER
V
248
R
-
ILE
A
312
L
H
ASN
V
109
R
-
ILE
A
312
L
H
PHE
V
110
R
-
ILE
A
312
L
H
ALA
V
247
R
-
PRO
A
313
L
H
ALA
V
247
R
-
PRO
A
313
L
-
ARG
V
250
R
-
PRO
A
313
L
-
LYS
V
252
R
-
GLU
A
314
L
-
LYS
V
252
R
-
GLU
A
314
L
-
ARG
V
253
R
-
GLU
A
314
L
-
GLU
V
254
R
-
GLY
A
315
L
-
ARG
V
253
R
H
PRO
A
316
L
H
LEU
V
104
R
H
PRO
A
316
L
-
SER
V
108
R
H
ALA
A
317
L
-
SER
V
108
R
H
ALA
A
317
L
H
PHE
V
110
R
H
TYR
A
320
L
H
LEU
V
104
R
H
TYR
A
320
L
H
PHE
V
110
R
-
HIS
A
448
L
-
GLU
V
107
R
-
HIS
A
450
L
-
LYS
V
258
R
H
PHE
A
456
L
H
GLU
V
103
R
H
PHE
A
456
L
-
GLY
V
105
R
H
LEU
A
459
L
-
GLY
V
106
R
H
VAL
A
460
L
H
LYS
V
102
R
H
VAL
A
460
L
-
GLY
V
105
R
H
VAL
A
460
L
-
GLY
V
106
R
H
ALA
A
463
L
-
GLY
V
106
R
H
LEU
A
464
L
H
LYS
V
102
R
H
LEU
A
464
L
H
LEU
V
115
R
H
THR
A
467
L
H
ASP
V
112
R
H
LEU
A
471
L
H
LYS
V
239
R
H
LEU
A
471
L
H
ARG
V
242
R
H
PHE
A
474
L
H
ARG
V
242
R
H
GLN
A
475
L
H
LYS
V
239
R
H
GLN
A
497
L
H
ARG
V
227
R
-
ASN
A
516
L
H
GLN
V
231
R
-
SER
A
518
L
H
ARG
V
123
R
-
SER
A
518
L
H
GLU
V
235
R
-
GLY
A
519
L
H
ARG
V
123
R
-
GLY
A
519
L
H
GLU
V
126
R
-
THR
A
520
L
H
ARG
V
123
R
-
THR
A
520
L
H
GLU
V
126
R
-
THR
A
522
L
-
VAL
V
81
R
-
LEU
A
523
L
H
ILE
V
133
R
-
LEU
A
525
L
H
GLN
V
228
R
-
THR
A
526
L
H
GLN
V
228
R
-
THR
A
526
L
H
GLN
V
231
R
-
THR
A
526
L
H
ILE
V
232
R
-
LEU
A
527
L
H
GLN
V
231
R
-
PRO
A
528
L
H
GLN
V
231
R
-
ARG
A
529
L
H
GLN
V
231
R
-
GLN
A
531
L
H
ARG
V
227
R
-
THR
A
541
L
-
ARG
V
78
R
-
THR
A
541
L
-
GLY
V
79
R
-
LYS
A
542
L
-
GLY
V
79
R
-
LYS
A
542
L
-
GLN
V
80
R
-
THR
A
552
L
H
ASP
V
224
R
-
GLY
A
553
L
H
ASP
V
220
R
-
GLY
A
554
L
H
GLU
V
134
R
-
GLY
A
554
L
H
ASP
V
220
R
-
GLY
A
554
L
H
ALA
V
221
R
-
GLY
A
554
L
H
ASP
V
224
R
-
HIS
A
555
L
H
LEU
V
131
R
-
HIS
A
555
L
H
GLU
V
134
R
-
HIS
A
555
L
H
ALA
V
221
R
-
HIS
A
555
L
H
ASP
V
224
R
-
HIS
A
555
L
H
TYR
V
225
R
-
HIS
A
555
L
H
GLN
V
228
A1IGU5_Q99961.1
contacts (residues & atoms)
Receptor
R: Secondary Structure
R: Atom
R: Residue
R; Chain
R: Residue number
Ligand
L: Secondary Structure
L; Atom
L: Residue
L: Chain
L: Residue Number
R
H
CE2
TRP
A
278
L
-
OE1
GLU
V
329
R
H
CH2
TRP
A
278
L
-
OE1
GLU
V
329
R
H
CZ2
TRP
A
278
L
-
CD
GLU
V
329
R
H
CZ2
TRP
A
278
L
-
OE1
GLU
V
329
R
H
C
LEU
A
281
L
-
HE1
TYR
V
315
R
H
CB
LEU
A
281
L
-
OH
TYR
V
315
R
H
CB
LEU
A
281
L
-
HH
TYR
V
315
R
H
O
LEU
A
281
L
-
HE1
TYR
V
315
R
H
N
CYS
A
282
L
-
HE1
TYR
V
315
R
H
CA
CYS
A
282
L
-
HE1
TYR
V
315
R
H
CB
GLU
A
285
L
-
CD1
TYR
V
315
R
H
CB
GLU
A
285
L
-
CE1
TYR
V
315
R
H
CB
GLU
A
285
L
-
OH
TYR
V
315
R
H
CB
GLU
A
285
L
-
CZ
TYR
V
315
R
H
CB
GLU
A
285
L
-
HD1
TYR
V
315
R
H
CB
GLU
A
285
L
-
HE1
TYR
V
315
R
H
CG
GLU
A
285
L
-
CG
TYR
V
315
R
H
CG
GLU
A
285
L
-
CD1
TYR
V
315
R
H
CG
GLU
A
285
L
-
CD2
TYR
V
315
R
H
CG
GLU
A
285
L
-
CE1
TYR
V
315
R
H
CG
GLU
A
285
L
-
CE2
TYR
V
315
R
H
CG
GLU
A
285
L
-
CZ
TYR
V
315
R
H
CG
GLU
A
285
L
-
HB3
TYR
V
315
R
H
CG
GLU
A
285
L
-
HD1
TYR
V
315
R
H
CG
GLU
A
285
L
-
HE1
TYR
V
315
R
H
CD
GLU
A
285
L
-
CG
TYR
V
315
R
H
CD
GLU
A
285
L
-
CD1
TYR
V
315
R
H
CD
GLU
A
285
L
-
CD2
TYR
V
315
R
H
CD
GLU
A
285
L
-
CE1
TYR
V
315
R
H
CD
GLU
A
285
L
-
CE2
TYR
V
315
R
H
CD
GLU
A
285
L
-
CZ
TYR
V
315
R
H
OE1
GLU
A
285
L
H
OH
TYR
V
359
R
H
OE1
GLU
A
285
L
H
CZ
TYR
V
359
R
H
OE2
GLU
A
285
L
-
CE2
TYR
V
315
R
H
OE2
GLU
A
285
L
-
CZ
TYR
V
315
R
H
CG
ASN
A
289
L
H
OH
TYR
V
359
R
H
ND2
ASN
A
289
L
H
OH
TYR
V
359
R
H
OD1
ASN
A
289
L
H
OH
TYR
V
359
R
H
OD1
ASN
A
289
L
H
HH
TYR
V
359
R
-
CA
LEU
A
310
L
H
HG3
MET
V
244
R
-
C
LEU
A
310
L
H
HA
MET
V
244
R
-
C
LEU
A
310
L
H
HG3
MET
V
244
R
-
CB
LEU
A
310
L
H
HD21
ASN
V
109
R
-
O
LEU
A
310
L
H
ND2
ASN
V
109
R
-
O
LEU
A
310
L
H
HD21
ASN
V
109
R
-
O
LEU
A
310
L
H
HD22
ASN
V
109
R
-
O
LEU
A
310
L
H
CA
MET
V
244
R
-
O
LEU
A
310
L
H
CG
MET
V
244
R
-
O
LEU
A
310
L
H
HA
MET
V
244
R
-
O
LEU
A
310
L
H
HG3
MET
V
244
R
-
O
LEU
A
310
L
H
CB
ALA
V
247
R
-
O
LEU
A
310
L
H
HB1
ALA
V
247
R
-
O
LEU
A
310
L
H
HB2
ALA
V
247
R
-
CG
LEU
A
310
L
H
HD2
PHE
V
110
R
-
CD1
LEU
A
310
L
H
CB
PHE
V
110
R
-
CD1
LEU
A
310
L
H
CG
PHE
V
110
R
-
CD1
LEU
A
310
L
H
CD2
PHE
V
110
R
-
CD1
LEU
A
310
L
H
HA
PHE
V
110
R
-
CD1
LEU
A
310
L
H
HB2
PHE
V
110
R
-
CD1
LEU
A
310
L
H
HD2
PHE
V
110
R
-
N
ASP
A
311
L
H
HG3
MET
V
244
R
-
CA
ASP
A
311
L
H
HG3
MET
V
244
R
-
CA
ASP
A
311
L
H
OG
SER
V
248
R
-
CA
ASP
A
311
L
H
HG
SER
V
248
R
-
C
ASP
A
311
L
H
C
ALA
V
247
R
-
C
ASP
A
311
L
H
CB
ALA
V
247
R
-
C
ASP
A
311
L
H
O
ALA
V
247
R
-
C
ASP
A
311
L
H
HB1
ALA
V
247
R
-
C
ASP
A
311
L
H
HB2
ALA
V
247
R
-
C
ASP
A
311
L
H
HA
SER
V
248
R
-
CB
ASP
A
311
L
H
O
MET
V
244
R
-
CB
ASP
A
311
L
H
HG2
MET
V
244
R
-
CB
ASP
A
311
L
H
HG3
MET
V
244
R
-
CB
ASP
A
311
L
H
HB2
ALA
V
247
R
-
CB
ASP
A
311
L
H
N
SER
V
248
R
-
CB
ASP
A
311
L
H
CA
SER
V
248
R
-
CB
ASP
A
311
L
H
CB
SER
V
248
R
-
CB
ASP
A
311
L
H
OG
SER
V
248
R
-
CB
ASP
A
311
L
H
H
SER
V
248
R
-
CB
ASP
A
311
L
H
HA
SER
V
248
R
-
CB
ASP
A
311
L
H
HB3
SER
V
248
R
-
CB
ASP
A
311
L
H
HG
SER
V
248
R
-
O
ASP
A
311
L
H
CA
ALA
V
247
R
-
O
ASP
A
311
L
H
C
ALA
V
247
R
-
O
ASP
A
311
L
H
CB
ALA
V
247
R
-
O
ASP
A
311
L
H
O
ALA
V
247
R
-
O
ASP
A
311
L
H
HA
ALA
V
247
R
-
O
ASP
A
311
L
H
HB1
ALA
V
247
R
-
O
ASP
A
311
L
H
HB2
ALA
V
247
R
-
O
ASP
A
311
L
H
N
SER
V
248
R
-
O
ASP
A
311
L
H
CA
SER
V
248
R
-
O
ASP
A
311
L
H
OG
SER
V
248
R
-
O
ASP
A
311
L
H
H
SER
V
248
R
-
O
ASP
A
311
L
H
HA
SER
V
248
R
-
CG
ASP
A
311
L
H
CG
MET
V
244
R
-
CG
ASP
A
311
L
H
HG2
MET
V
244
R
-
CG
ASP
A
311
L
H
HG3
MET
V
244
R
-
CG
ASP
A
311
L
H
CB
SER
V
248
R
-
CG
ASP
A
311
L
H
OG
SER
V
248
R
-
CG
ASP
A
311
L
H
HG
SER
V
248
R
-
OD1
ASP
A
311
L
H
OG
SER
V
248
R
-
OD2
ASP
A
311
L
H
CG
MET
V
244
R
-
OD2
ASP
A
311
L
H
SD
MET
V
244
R
-
OD2
ASP
A
311
L
H
HG2
MET
V
244
R
-
OD2
ASP
A
311
L
H
HG3
MET
V
244
R
-
OD2
ASP
A
311
L
H
OG
SER
V
248
R
-
OD2
ASP
A
311
L
H
HG
SER
V
248
R
-
N
ILE
A
312
L
H
HB1
ALA
V
247
R
-
N
ILE
A
312
L
H
HB2
ALA
V
247
R
-
CA
ILE
A
312
L
H
HB1
ALA
V
247
R
-
CB
ILE
A
312
L
H
HB3
ASN
V
109
R
-
CB
ILE
A
312
L
H
HD21
ASN
V
109
R
-
CB
ILE
A
312
L
H
HB1
ALA
V
247
R
-
CG1
ILE
A
312
L
H
CB
ASN
V
109
R
-
CG1
ILE
A
312
L
H
CG
ASN
V
109
R
-
CG1
ILE
A
312
L
H
ND2
ASN
V
109
R
-
CG1
ILE
A
312
L
H
HB2
ASN
V
109
R
-
CG1
ILE
A
312
L
H
HB3
ASN
V
109
R
-
CG1
ILE
A
312
L
H
HD21
ASN
V
109
R
-
CG1
ILE
A
312
L
H
HD22
ASN
V
109
R
-
CG1
ILE
A
312
L
H
CB
ALA
V
247
R
-
CG1
ILE
A
312
L
H
HB1
ALA
V
247
R
-
CG1
ILE
A
312
L
H
HB2
ALA
V
247
R
-
CG1
ILE
A
312
L
H
HB3
ALA
V
247
R
-
CG2
ILE
A
312
L
H
CB
ASN
V
109
R
-
CG2
ILE
A
312
L
H
HB2
ASN
V
109
R
-
CG2
ILE
A
312
L
H
HB3
ASN
V
109
R
-
CD1
ILE
A
312
L
H
CA
ASN
V
109
R
-
CD1
ILE
A
312
L
H
C
ASN
V
109
R
-
CD1
ILE
A
312
L
H
CB
ASN
V
109
R
-
CD1
ILE
A
312
L
H
O
ASN
V
109
R
-
CD1
ILE
A
312
L
H
CG
ASN
V
109
R
-
CD1
ILE
A
312
L
H
ND2
ASN
V
109
R
-
CD1
ILE
A
312
L
H
OD1
ASN
V
109
R
-
CD1
ILE
A
312
L
H
H
ASN
V
109
R
-
CD1
ILE
A
312
L
H
HB2
ASN
V
109
R
-
CD1
ILE
A
312
L
H
HB3
ASN
V
109
R
-
CD1
ILE
A
312
L
H
HD21
ASN
V
109
R
-
CD1
ILE
A
312
L
H
HD22
ASN
V
109
R
-
CD1
ILE
A
312
L
H
N
PHE
V
110
R
-
CD1
ILE
A
312
L
H
H
PHE
V
110
R
-
CD1
ILE
A
312
L
H
HA
PHE
V
110
R
-
CD1
ILE
A
312
L
H
HB2
PHE
V
110
R
-
CA
PRO
A
313
L
-
HG2
LYS
V
252
R
-
CA
PRO
A
313
L
-
HE2
LYS
V
252
R
-
C
PRO
A
313
L
-
HG2
LYS
V
252
R
-
CB
PRO
A
313
L
-
CG
LYS
V
252
R
-
CB
PRO
A
313
L
-
CE
LYS
V
252
R
-
CB
PRO
A
313
L
-
HA
LYS
V
252
R
-
CB
PRO
A
313
L
-
HG2
LYS
V
252
R
-
CB
PRO
A
313
L
-
HG3
LYS
V
252
R
-
CB
PRO
A
313
L
-
HE2
LYS
V
252
R
-
O
PRO
A
313
L
-
HG2
LYS
V
252
R
-
CG
PRO
A
313
L
-
O
ARG
V
250
R
-
CG
PRO
A
313
L
-
HA
LYS
V
252
R
-
CG
PRO
A
313
L
-
HG2
LYS
V
252
R
-
CG
PRO
A
313
L
-
HG3
LYS
V
252
R
-
CD
PRO
A
313
L
H
O
ALA
V
247
R
-
CB
GLU
A
314
L
-
HA
LYS
V
252
R
-
CB
GLU
A
314
L
-
N
ARG
V
253
R
-
CB
GLU
A
314
L
-
CB
ARG
V
253
R
-
CB
GLU
A
314
L
-
O
ARG
V
253
R
-
CB
GLU
A
314
L
-
H
ARG
V
253
R
-
CB
GLU
A
314
L
-
HB2
ARG
V
253
R
-
CB
GLU
A
314
L
-
HG3
ARG
V
253
R
-
CG
GLU
A
314
L
-
CA
LYS
V
252
R
-
CG
GLU
A
314
L
-
C
LYS
V
252
R
-
CG
GLU
A
314
L
-
CB
LYS
V
252
R
-
CG
GLU
A
314
L
-
HA
LYS
V
252
R
-
CG
GLU
A
314
L
-
HB3
LYS
V
252
R
-
CG
GLU
A
314
L
-
HG2
LYS
V
252
R
-
CG
GLU
A
314
L
-
N
ARG
V
253
R
-
CG
GLU
A
314
L
-
CA
ARG
V
253
R
-
CG
GLU
A
314
L
-
C
ARG
V
253
R
-
CG
GLU
A
314
L
-
O
ARG
V
253
R
-
CG
GLU
A
314
L
-
H
ARG
V
253
R
-
CG
GLU
A
314
L
-
HB2
ARG
V
253
R
-
CD
GLU
A
314
L
-
CA
LYS
V
252
R
-
CD
GLU
A
314
L
-
C
LYS
V
252
R
-
CD
GLU
A
314
L
-
CB
LYS
V
252
R
-
CD
GLU
A
314
L
-
O
LYS
V
252
R
-
CD
GLU
A
314
L
-
HA
LYS
V
252
R
-
CD
GLU
A
314
L
-
HB3
LYS
V
252
R
-
CD
GLU
A
314
L
-
HG2
LYS
V
252
R
-
CD
GLU
A
314
L
-
N
ARG
V
253
R
-
CD
GLU
A
314
L
-
CA
ARG
V
253
R
-
CD
GLU
A
314
L
-
C
ARG
V
253
R
-
CD
GLU
A
314
L
-
CB
ARG
V
253
R
-
CD
GLU
A
314
L
-
O
ARG
V
253
R
-
CD
GLU
A
314
L
-
H
ARG
V
253
R
-
CD
GLU
A
314
L
-
HA
ARG
V
253
R
-
CD
GLU
A
314
L
-
HB2
ARG
V
253
R
-
CD
GLU
A
314
L
-
HB3
ARG
V
253
R
-
CD
GLU
A
314
L
-
N
GLU
V
254
R
-
OE1
GLU
A
314
L
-
CA
LYS
V
252
R
-
OE1
GLU
A
314
L
-
C
LYS
V
252
R
-
OE1
GLU
A
314
L
-
CB
LYS
V
252
R
-
OE1
GLU
A
314
L
-
O
LYS
V
252
R
-
OE1
GLU
A
314
L
-
CG
LYS
V
252
R
-
OE1
GLU
A
314
L
-
HA
LYS
V
252
R
-
OE1
GLU
A
314
L
-
HB2
LYS
V
252
R
-
OE1
GLU
A
314
L
-
HB3
LYS
V
252
R
-
OE1
GLU
A
314
L
-
HG2
LYS
V
252
R
-
OE1
GLU
A
314
L
-
HD3
LYS
V
252
R
-
OE1
GLU
A
314
L
-
N
ARG
V
253
R
-
OE1
GLU
A
314
L
-
CA
ARG
V
253
R
-
OE1
GLU
A
314
L
-
C
ARG
V
253
R
-
OE1
GLU
A
314
L
-
O
ARG
V
253
R
-
OE1
GLU
A
314
L
-
H
ARG
V
253
R
-
OE1
GLU
A
314
L
-
N
GLU
V
254
R
-
OE1
GLU
A
314
L
-
HA
GLU
V
254
R
-
OE2
GLU
A
314
L
-
CA
LYS
V
252
R
-
OE2
GLU
A
314
L
-
C
LYS
V
252
R
-
OE2
GLU
A
314
L
-
O
LYS
V
252
R
-
OE2
GLU
A
314
L
-
HA
LYS
V
252
R
-
OE2
GLU
A
314
L
-
HB3
LYS
V
252
R
-
OE2
GLU
A
314
L
-
N
ARG
V
253
R
-
OE2
GLU
A
314
L
-
CA
ARG
V
253
R
-
OE2
GLU
A
314
L
-
C
ARG
V
253
R
-
OE2
GLU
A
314
L
-
CB
ARG
V
253
R
-
OE2
GLU
A
314
L
-
O
ARG
V
253
R
-
OE2
GLU
A
314
L
-
CG
ARG
V
253
R
-
OE2
GLU
A
314
L
-
H
ARG
V
253
R
-
OE2
GLU
A
314
L
-
HA
ARG
V
253
R
-
OE2
GLU
A
314
L
-
HB2
ARG
V
253
R
-
OE2
GLU
A
314
L
-
HB3
ARG
V
253
R
-
OE2
GLU
A
314
L
-
HG2
ARG
V
253
R
-
OE2
GLU
A
314
L
-
HG3
ARG
V
253
R
-
OE2
GLU
A
314
L
-
N
GLU
V
254
R
-
OE2
GLU
A
314
L
-
H
GLU
V
254
R
-
CA
GLY
A
315
L
-
HD2
ARG
V
253
R
H
C
PRO
A
316
L
-
HG
SER
V
108
R
H
CB
PRO
A
316
L
-
HB3
SER
V
108
R
H
O
PRO
A
316
L
H
HD22
LEU
V
104
R
H
CG
PRO
A
316
L
-
HB3
SER
V
108
R
H
N
ALA
A
317
L
-
OG
SER
V
108
R
H
N
ALA
A
317
L
-
HG
SER
V
108
R
H
CA
ALA
A
317
L
-
OG
SER
V
108
R
H
CA
ALA
A
317
L
-
HG
SER
V
108
R
H
CA
ALA
A
317
L
H
HB3
PHE
V
110
R
H
CB
ALA
A
317
L
-
OG
SER
V
108
R
H
CB
ALA
A
317
L
-
HG
SER
V
108
R
H
CB
ALA
A
317
L
H
CB
PHE
V
110
R
H
CB
ALA
A
317
L
H
H
PHE
V
110
R
H
CB
ALA
A
317
L
H
HB2
PHE
V
110
R
H
CB
ALA
A
317
L
H
HB3
PHE
V
110
R
H
CB
TYR
A
320
L
H
HD22
LEU
V
104
R
H
CB
TYR
A
320
L
H
HD23
LEU
V
104
R
H
CG
TYR
A
320
L
H
HD12
LEU
V
104
R
H
CG
TYR
A
320
L
H
HD23
LEU
V
104
R
H
CG
TYR
A
320
L
H
HD1
PHE
V
110
R
H
CG
TYR
A
320
L
H
HE1
PHE
V
110
R
H
CD1
TYR
A
320
L
H
CD1
PHE
V
110
R
H
CD1
TYR
A
320
L
H
CE1
PHE
V
110
R
H
CD1
TYR
A
320
L
H
HD1
PHE
V
110
R
H
CD1
TYR
A
320
L
H
HE1
PHE
V
110
R
H
CD2
TYR
A
320
L
H
HD23
LEU
V
104
R
H
CE1
TYR
A
320
L
H
CD1
PHE
V
110
R
H
CE1
TYR
A
320
L
H
CE1
PHE
V
110
R
H
CE1
TYR
A
320
L
H
CZ
PHE
V
110
R
H
CE1
TYR
A
320
L
H
HD1
PHE
V
110
R
H
CE1
TYR
A
320
L
H
HE1
PHE
V
110
R
H
CE1
TYR
A
320
L
H
HZ
PHE
V
110
R
H
CE2
TYR
A
320
L
H
HD23
LEU
V
104
R
H
CE2
TYR
A
320
L
H
HE1
PHE
V
110
R
H
OH
TYR
A
320
L
H
HE1
PHE
V
110
R
H
CZ
TYR
A
320
L
H
CE1
PHE
V
110
R
H
CZ
TYR
A
320
L
H
HE1
PHE
V
110
R
-
CE1
HIS
A
448
L
-
CD
GLU
V
107
R
-
CE1
HIS
A
448
L
-
OE1
GLU
V
107
R
-
CE1
HIS
A
448
L
-
HB2
GLU
V
107
R
-
CE1
HIS
A
448
L
-
HG2
GLU
V
107
R
-
NE2
HIS
A
448
L
-
CB
GLU
V
107
R
-
NE2
HIS
A
448
L
-
CG
GLU
V
107
R
-
NE2
HIS
A
448
L
-
CD
GLU
V
107
R
-
NE2
HIS
A
448
L
-
OE1
GLU
V
107
R
-
NE2
HIS
A
448
L
-
HB2
GLU
V
107
R
-
NE2
HIS
A
448
L
-
HG2
GLU
V
107
R
-
CD2
HIS
A
450
L
-
NZ
LYS
V
258
R
-
CD2
HIS
A
450
L
-
HZ1
LYS
V
258
R
-
CD2
HIS
A
450
L
-
HZ2
LYS
V
258
R
-
CD2
HIS
A
450
L
-
HZ3
LYS
V
258
R
-
CE1
HIS
A
450
L
-
HZ2
LYS
V
258
R
-
NE2
HIS
A
450
L
-
NZ
LYS
V
258
R
-
NE2
HIS
A
450
L
-
HZ1
LYS
V
258
R
-
NE2
HIS
A
450
L
-
HZ2
LYS
V
258
R
-
NE2
HIS
A
450
L
-
HZ3
LYS
V
258
R
H
CE1
PHE
A
456
L
H
O
GLU
V
103
R
H
CE1
PHE
A
456
L
H
HA
GLU
V
103
R
H
CE2
PHE
A
456
L
-
HA3
GLY
V
105
R
H
CZ
PHE
A
456
L
H
C
GLU
V
103
R
H
CZ
PHE
A
456
L
H
O
GLU
V
103
R
H
CZ
PHE
A
456
L
H
HA
GLU
V
103
R
H
CZ
PHE
A
456
L
-
HA2
GLY
V
105
R
H
CZ
PHE
A
456
L
-
HA3
GLY
V
105
R
H
CD2
LEU
A
459
L
-
H
GLY
V
106
R
H
CA
VAL
A
460
L
-
H
GLY
V
106
R
H
CB
VAL
A
460
L
H
O
LYS
V
102
R
H
CB
VAL
A
460
L
-
HA2
GLY
V
105
R
H
CG1
VAL
A
460
L
H
CA
LYS
V
102
R
H
CG1
VAL
A
460
L
H
O
LYS
V
102
R
H
CG1
VAL
A
460
L
H
HA
LYS
V
102
R
H
CG1
VAL
A
460
L
H
HB2
LYS
V
102
R
H
CG1
VAL
A
460
L
H
HG2
LYS
V
102
R
H
CG1
VAL
A
460
L
-
HA2
GLY
V
105
R
H
CG2
VAL
A
460
L
H
O
LYS
V
102
R
H
CG2
VAL
A
460
L
-
CA
GLY
V
105
R
H
CG2
VAL
A
460
L
-
HA2
GLY
V
105
R
H
CG2
VAL
A
460
L
-
HA3
GLY
V
105
R
H
CG2
VAL
A
460
L
-
H
GLY
V
106
R
H
CA
ALA
A
463
L
-
HA3
GLY
V
106
R
H
CB
ALA
A
463
L
-
CA
GLY
V
106
R
H
CB
ALA
A
463
L
-
H
GLY
V
106
R
H
CB
ALA
A
463
L
-
HA2
GLY
V
106
R
H
CB
ALA
A
463
L
-
HA3
GLY
V
106
R
H
CB
LEU
A
464
L
H
HD12
LEU
V
115
R
H
CG
LEU
A
464
L
H
CD1
LEU
V
115
R
H
CG
LEU
A
464
L
H
HD11
LEU
V
115
R
H
CG
LEU
A
464
L
H
HD12
LEU
V
115
R
H
CG
LEU
A
464
L
H
HD13
LEU
V
115
R
H
CD1
LEU
A
464
L
H
HG2
LYS
V
102
R
H
CD1
LEU
A
464
L
H
HE2
LYS
V
102
R
H
CD1
LEU
A
464
L
H
CD1
LEU
V
115
R
H
CD1
LEU
A
464
L
H
HD12
LEU
V
115
R
H
CD1
LEU
A
464
L
H
HD13
LEU
V
115
R
H
CD1
LEU
A
464
L
H
HD22
LEU
V
115
R
H
CD2
LEU
A
464
L
H
CG
LEU
V
115
R
H
CD2
LEU
A
464
L
H
CD1
LEU
V
115
R
H
CD2
LEU
A
464
L
H
HD11
LEU
V
115
R
H
CD2
LEU
A
464
L
H
HD12
LEU
V
115
R
H
CD2
LEU
A
464
L
H
HD13
LEU
V
115
R
H
CD2
LEU
A
464
L
H
HD22
LEU
V
115
R
H
CG2
THR
A
467
L
H
OD2
ASP
V
112
R
H
CG
LEU
A
471
L
H
HE2
LYS
V
239
R
H
CG
LEU
A
471
L
H
HH22
ARG
V
242
R
H
CD1
LEU
A
471
L
H
HG2
LYS
V
239
R
H
CD1
LEU
A
471
L
H
HE2
LYS
V
239
R
H
CD1
LEU
A
471
L
H
NH2
ARG
V
242
R
H
CD1
LEU
A
471
L
H
CZ
ARG
V
242
R
H
CD1
LEU
A
471
L
H
HH21
ARG
V
242
R
H
CD1
LEU
A
471
L
H
HH22
ARG
V
242
R
H
CD2
LEU
A
471
L
H
CE
LYS
V
239
R
H
CD2
LEU
A
471
L
H
HE2
LYS
V
239
R
H
CD2
LEU
A
471
L
H
HZ3
LYS
V
239
R
H
CD1
PHE
A
474
L
H
NH1
ARG
V
242
R
H
CD1
PHE
A
474
L
H
HH11
ARG
V
242
R
H
CE1
PHE
A
474
L
H
NH1
ARG
V
242
R
H
CE1
PHE
A
474
L
H
HD3
ARG
V
242
R
H
CE1
PHE
A
474
L
H
HH11
ARG
V
242
R
H
CE1
PHE
A
474
L
H
HH12
ARG
V
242
R
H
CZ
PHE
A
474
L
H
HD2
ARG
V
242
R
H
CZ
PHE
A
474
L
H
HD3
ARG
V
242
R
H
CZ
PHE
A
474
L
H
HH11
ARG
V
242
R
H
NE2
GLN
A
475
L
H
HE2
LYS
V
239
R
H
CB
GLN
A
497
L
H
HH12
ARG
V
227
R
H
CB
GLN
A
497
L
H
HH22
ARG
V
227
R
H
CG
GLN
A
497
L
H
NH1
ARG
V
227
R
H
CG
GLN
A
497
L
H
HH12
ARG
V
227
R
H
CG
GLN
A
497
L
H
HH22
ARG
V
227
R
H
CD
GLN
A
497
L
H
NH1
ARG
V
227
R
H
CD
GLN
A
497
L
H
NH2
ARG
V
227
R
H
CD
GLN
A
497
L
H
CZ
ARG
V
227
R
H
CD
GLN
A
497
L
H
HH11
ARG
V
227
R
H
CD
GLN
A
497
L
H
HH12
ARG
V
227
R
H
CD
GLN
A
497
L
H
HH22
ARG
V
227
R
H
NE2
GLN
A
497
L
H
NH1
ARG
V
227
R
H
NE2
GLN
A
497
L
H
CZ
ARG
V
227
R
H
NE2
GLN
A
497
L
H
HH11
ARG
V
227
R
H
NE2
GLN
A
497
L
H
HH12
ARG
V
227
R
H
NE2
GLN
A
497
L
H
HH22
ARG
V
227
R
H
OE1
GLN
A
497
L
H
NH1
ARG
V
227
R
H
OE1
GLN
A
497
L
H
NH2
ARG
V
227
R
H
OE1
GLN
A
497
L
H
CZ
ARG
V
227
R
H
OE1
GLN
A
497
L
H
HH11
ARG
V
227
R
H
OE1
GLN
A
497
L
H
HH12
ARG
V
227
R
H
OE1
GLN
A
497
L
H
HH21
ARG
V
227
R
H
OE1
GLN
A
497
L
H
HH22
ARG
V
227
R
-
CG
ASN
A
516
L
H
HE21
GLN
V
231
R
-
OD1
ASN
A
516
L
H
HE21
GLN
V
231
R
-
CB
SER
A
518
L
H
NH1
ARG
V
123
R
-
CB
SER
A
518
L
H
HH11
ARG
V
123
R
-
CB
SER
A
518
L
H
HH12
ARG
V
123
R
-
OG
SER
A
518
L
H
NH1
ARG
V
123
R
-
OG
SER
A
518
L
H
HH11
ARG
V
123
R
-
OG
SER
A
518
L
H
HH12
ARG
V
123
R
-
OG
SER
A
518
L
H
CD
GLU
V
235
R
-
OG
SER
A
518
L
H
OE1
GLU
V
235
R
-
OG
SER
A
518
L
H
OE2
GLU
V
235
R
-
O
GLY
A
519
L
H
CG
ARG
V
123
R
-
O
GLY
A
519
L
H
CD
ARG
V
123
R
-
O
GLY
A
519
L
H
HG2
ARG
V
123
R
-
O
GLY
A
519
L
H
HD3
ARG
V
123
R
-
O
GLY
A
519
L
H
OE2
GLU
V
126
R
-
O
THR
A
520
L
H
OE1
GLU
V
126
R
-
O
THR
A
520
L
H
OE2
GLU
V
126
R
-
CG2
THR
A
520
L
H
NH2
ARG
V
123
R
-
CG2
THR
A
520
L
H
HH21
ARG
V
123
R
-
CG2
THR
A
522
L
-
O
VAL
V
81
R
-
CD2
LEU
A
523
L
H
HD12
ILE
V
133
R
-
CA
LEU
A
525
L
H
OE1
GLN
V
228
R
-
C
LEU
A
525
L
H
OE1
GLN
V
228
R
-
CD2
LEU
A
525
L
H
CD
GLN
V
228
R
-
CD2
LEU
A
525
L
H
NE2
GLN
V
228
R
-
CD2
LEU
A
525
L
H
OE1
GLN
V
228
R
-
CD2
LEU
A
525
L
H
HE22
GLN
V
228
R
-
N
THR
A
526
L
H
OE1
GLN
V
228
R
-
N
THR
A
526
L
H
HG2
GLN
V
228
R
-
CA
THR
A
526
L
H
OE1
GLN
V
231
R
-
C
THR
A
526
L
H
HG2
GLN
V
228
R
-
C
THR
A
526
L
H
CD
GLN
V
231
R
-
C
THR
A
526
L
H
OE1
GLN
V
231
R
-
CB
THR
A
526
L
H
HB2
GLN
V
228
R
-
CB
THR
A
526
L
H
OE1
GLN
V
231
R
-
CB
THR
A
526
L
H
HG12
ILE
V
232
R
-
CB
THR
A
526
L
H
HD11
ILE
V
232
R
-
O
THR
A
526
L
H
CG
GLN
V
228
R
-
O
THR
A
526
L
H
HA
GLN
V
228
R
-
O
THR
A
526
L
H
HB2
GLN
V
228
R
-
O
THR
A
526
L
H
HG2
GLN
V
228
R
-
O
THR
A
526
L
H
CD
GLN
V
231
R
-
O
THR
A
526
L
H
OE1
GLN
V
231
R
-
CG2
THR
A
526
L
H
CB
GLN
V
231
R
-
CG2
THR
A
526
L
H
CG
GLN
V
231
R
-
CG2
THR
A
526
L
H
CD
GLN
V
231
R
-
CG2
THR
A
526
L
H
OE1
GLN
V
231
R
-
CG2
THR
A
526
L
H
HB2
GLN
V
231
R
-
CG2
THR
A
526
L
H
HG2
GLN
V
231
R
-
CG2
THR
A
526
L
H
CG1
ILE
V
232
R
-
CG2
THR
A
526
L
H
HG12
ILE
V
232
R
-
CG2
THR
A
526
L
H
HG13
ILE
V
232
R
-
OG1
THR
A
526
L
H
HG12
ILE
V
232
R
-
OG1
THR
A
526
L
H
HG13
ILE
V
232
R
-
OG1
THR
A
526
L
H
HD11
ILE
V
232
R
-
N
LEU
A
527
L
H
CD
GLN
V
231
R
-
N
LEU
A
527
L
H
NE2
GLN
V
231
R
-
N
LEU
A
527
L
H
OE1
GLN
V
231
R
-
N
LEU
A
527
L
H
HE22
GLN
V
231
R
-
CA
LEU
A
527
L
H
CD
GLN
V
231
R
-
CA
LEU
A
527
L
H
NE2
GLN
V
231
R
-
CA
LEU
A
527
L
H
OE1
GLN
V
231
R
-
CA
LEU
A
527
L
H
HE22
GLN
V
231
R
-
C
LEU
A
527
L
H
CD
GLN
V
231
R
-
C
LEU
A
527
L
H
NE2
GLN
V
231
R
-
C
LEU
A
527
L
H
OE1
GLN
V
231
R
-
C
LEU
A
527
L
H
HE21
GLN
V
231
R
-
C
LEU
A
527
L
H
HE22
GLN
V
231
R
-
CB
LEU
A
527
L
H
HE22
GLN
V
231
R
-
O
LEU
A
527
L
H
CD
GLN
V
231
R
-
O
LEU
A
527
L
H
NE2
GLN
V
231
R
-
O
LEU
A
527
L
H
OE1
GLN
V
231
R
-
O
LEU
A
527
L
H
HE21
GLN
V
231
R
-
O
LEU
A
527
L
H
HE22
GLN
V
231
R
-
N
PRO
A
528
L
H
CD
GLN
V
231
R
-
N
PRO
A
528
L
H
NE2
GLN
V
231
R
-
N
PRO
A
528
L
H
OE1
GLN
V
231
R
-
N
PRO
A
528
L
H
HE21
GLN
V
231
R
-
N
PRO
A
528
L
H
HE22
GLN
V
231
R
-
CA
PRO
A
528
L
H
CD
GLN
V
231
R
-
CA
PRO
A
528
L
H
NE2
GLN
V
231
R
-
CA
PRO
A
528
L
H
OE1
GLN
V
231
R
-
CA
PRO
A
528
L
H
HE21
GLN
V
231
R
-
CA
PRO
A
528
L
H
HE22
GLN
V
231
R
-
C
PRO
A
528
L
H
NE2
GLN
V
231
R
-
C
PRO
A
528
L
H
HE21
GLN
V
231
R
-
C
PRO
A
528
L
H
HE22
GLN
V
231
R
-
CB
PRO
A
528
L
H
CD
GLN
V
231
R
-
CB
PRO
A
528
L
H
NE2
GLN
V
231
R
-
CB
PRO
A
528
L
H
HG3
GLN
V
231
R
-
CB
PRO
A
528
L
H
HE21
GLN
V
231
R
-
CB
PRO
A
528
L
H
HE22
GLN
V
231
R
-
O
PRO
A
528
L
H
HE22
GLN
V
231
R
-
CG
PRO
A
528
L
H
CD
GLN
V
231
R
-
CG
PRO
A
528
L
H
NE2
GLN
V
231
R
-
CG
PRO
A
528
L
H
HE21
GLN
V
231
R
-
CG
PRO
A
528
L
H
HE22
GLN
V
231
R
-
CD
PRO
A
528
L
H
CG
GLN
V
231
R
-
CD
PRO
A
528
L
H
CD
GLN
V
231
R
-
CD
PRO
A
528
L
H
NE2
GLN
V
231
R
-
CD
PRO
A
528
L
H
OE1
GLN
V
231
R
-
CD
PRO
A
528
L
H
HE21
GLN
V
231
R
-
CD
PRO
A
528
L
H
HE22
GLN
V
231
R
-
N
ARG
A
529
L
H
HE22
GLN
V
231
R
-
NE2
GLN
A
531
L
H
HD3
ARG
V
227
R
-
CB
THR
A
541
L
-
HA2
GLY
V
79
R
-
O
THR
A
541
L
-
HE
ARG
V
78
R
-
OG1
THR
A
541
L
-
CA
GLY
V
79
R
-
OG1
THR
A
541
L
-
HA2
GLY
V
79
R
-
OG1
THR
A
541
L
-
HA3
GLY
V
79
R
-
CE
LYS
A
542
L
-
O
GLY
V
79
R
-
CE
LYS
A
542
L
-
CG
GLN
V
80
R
-
CE
LYS
A
542
L
-
HG2
GLN
V
80
R
-
CE
LYS
A
542
L
-
HG3
GLN
V
80
R
-
CE
LYS
A
542
L
-
HE21
GLN
V
80
R
-
NZ
LYS
A
542
L
-
O
GLY
V
79
R
-
NZ
LYS
A
542
L
-
CG
GLN
V
80
R
-
NZ
LYS
A
542
L
-
CD
GLN
V
80
R
-
NZ
LYS
A
542
L
-
NE2
GLN
V
80
R
-
NZ
LYS
A
542
L
-
HG2
GLN
V
80
R
-
NZ
LYS
A
542
L
-
HG3
GLN
V
80
R
-
NZ
LYS
A
542
L
-
HE21
GLN
V
80
R
-
NZ
LYS
A
542
L
-
HE22
GLN
V
80
R
-
CB
THR
A
552
L
H
OD1
ASP
V
224
R
-
OG1
THR
A
552
L
H
CG
ASP
V
224
R
-
OG1
THR
A
552
L
H
OD1
ASP
V
224
R
-
C
GLY
A
553
L
H
HB3
ASP
V
220
R
-
O
GLY
A
553
L
H
CB
ASP
V
220
R
-
O
GLY
A
553
L
H
CG
ASP
V
220
R
-
O
GLY
A
553
L
H
OD1
ASP
V
220
R
-
O
GLY
A
553
L
H
HB3
ASP
V
220
R
-
N
GLY
A
554
L
H
HB3
ASP
V
220
R
-
N
GLY
A
554
L
H
HB2
ASP
V
224
R
-
CA
GLY
A
554
L
H
C
ASP
V
220
R
-
CA
GLY
A
554
L
H
CB
ASP
V
220
R
-
CA
GLY
A
554
L
H
O
ASP
V
220
R
-
CA
GLY
A
554
L
H
HB3
ASP
V
220
R
-
CA
GLY
A
554
L
H
CB
ASP
V
224
R
-
CA
GLY
A
554
L
H
HB2
ASP
V
224
R
-
C
GLY
A
554
L
H
O
ASP
V
220
R
-
C
GLY
A
554
L
H
HB3
ASP
V
220
R
-
C
GLY
A
554
L
H
HA
ALA
V
221
R
-
C
GLY
A
554
L
H
CB
ASP
V
224
R
-
C
GLY
A
554
L
H
CG
ASP
V
224
R
-
C
GLY
A
554
L
H
HB2
ASP
V
224
R
-
C
GLY
A
554
L
H
HB3
ASP
V
224
R
-
O
GLY
A
554
L
H
HG2
GLU
V
134
R
-
O
GLY
A
554
L
H
HB3
ASP
V
220
R
-
O
GLY
A
554
L
H
HA
ALA
V
221
R
-
O
GLY
A
554
L
H
HB2
ASP
V
224
R
-
O
GLY
A
554
L
H
HB3
ASP
V
224
R
-
N
HIS
A
555
L
H
CB
ASP
V
224
R
-
N
HIS
A
555
L
H
CG
ASP
V
224
R
-
N
HIS
A
555
L
H
OD1
ASP
V
224
R
-
N
HIS
A
555
L
H
OD2
ASP
V
224
R
-
N
HIS
A
555
L
H
HB2
ASP
V
224
R
-
N
HIS
A
555
L
H
HB3
ASP
V
224
R
-
CA
HIS
A
555
L
H
HG2
GLU
V
134
R
-
CA
HIS
A
555
L
H
CB
ASP
V
224
R
-
CA
HIS
A
555
L
H
CG
ASP
V
224
R
-
CA
HIS
A
555
L
H
OD1
ASP
V
224
R
-
CA
HIS
A
555
L
H
OD2
ASP
V
224
R
-
CA
HIS
A
555
L
H
HB2
ASP
V
224
R
-
CA
HIS
A
555
L
H
HB3
ASP
V
224
R
-
CB
HIS
A
555
L
H
CB
ASP
V
224
R
-
CB
HIS
A
555
L
H
CG
ASP
V
224
R
-
CB
HIS
A
555
L
H
OD1
ASP
V
224
R
-
CB
HIS
A
555
L
H
OD2
ASP
V
224
R
-
CB
HIS
A
555
L
H
HB2
ASP
V
224
R
-
CB
HIS
A
555
L
H
HB3
ASP
V
224
R
-
CB
HIS
A
555
L
H
NE2
GLN
V
228
R
-
CB
HIS
A
555
L
H
HE21
GLN
V
228
R
-
CB
HIS
A
555
L
H
HE22
GLN
V
228
R
-
CG
HIS
A
555
L
H
HG2
GLU
V
134
R
-
CG
HIS
A
555
L
H
HA
ALA
V
221
R
-
CG
HIS
A
555
L
H
CA
ASP
V
224
R
-
CG
HIS
A
555
L
H
C
ASP
V
224
R
-
CG
HIS
A
555
L
H
CB
ASP
V
224
R
-
CG
HIS
A
555
L
H
CG
ASP
V
224
R
-
CG
HIS
A
555
L
H
OD1
ASP
V
224
R
-
CG
HIS
A
555
L
H
OD2
ASP
V
224
R
-
CG
HIS
A
555
L
H
HB2
ASP
V
224
R
-
CG
HIS
A
555
L
H
HB3
ASP
V
224
R
-
CG
HIS
A
555
L
H
N
TYR
V
225
R
-
CG
HIS
A
555
L
H
H
TYR
V
225
R
-
CG
HIS
A
555
L
H
NE2
GLN
V
228
R
-
CG
HIS
A
555
L
H
HE21
GLN
V
228
R
-
CG
HIS
A
555
L
H
HE22
GLN
V
228
R
-
CD2
HIS
A
555
L
H
O
ALA
V
221
R
-
CD2
HIS
A
555
L
H
N
ASP
V
224
R
-
CD2
HIS
A
555
L
H
CA
ASP
V
224
R
-
CD2
HIS
A
555
L
H
C
ASP
V
224
R
-
CD2
HIS
A
555
L
H
CB
ASP
V
224
R
-
CD2
HIS
A
555
L
H
O
ASP
V
224
R
-
CD2
HIS
A
555
L
H
CG
ASP
V
224
R
-
CD2
HIS
A
555
L
H
OD1
ASP
V
224
R
-
CD2
HIS
A
555
L
H
OD2
ASP
V
224
R
-
CD2
HIS
A
555
L
H
H
ASP
V
224
R
-
CD2
HIS
A
555
L
H
HA
ASP
V
224
R
-
CD2
HIS
A
555
L
H
HB2
ASP
V
224
R
-
CD2
HIS
A
555
L
H
HB3
ASP
V
224
R
-
CD2
HIS
A
555
L
H
N
TYR
V
225
R
-
CD2
HIS
A
555
L
H
CA
TYR
V
225
R
-
CD2
HIS
A
555
L
H
H
TYR
V
225
R
-
CD2
HIS
A
555
L
H
HA
TYR
V
225
R
-
CD2
HIS
A
555
L
H
NE2
GLN
V
228
R
-
CD2
HIS
A
555
L
H
HE21
GLN
V
228
R
-
CD2
HIS
A
555
L
H
HE22
GLN
V
228
R
-
ND1
HIS
A
555
L
H
HA
LEU
V
131
R
-
ND1
HIS
A
555
L
H
CG
GLU
V
134
R
-
ND1
HIS
A
555
L
H
HG2
GLU
V
134
R
-
ND1
HIS
A
555
L
H
CA
ALA
V
221
R
-
ND1
HIS
A
555
L
H
O
ALA
V
221
R
-
ND1
HIS
A
555
L
H
HA
ALA
V
221
R
-
ND1
HIS
A
555
L
H
CB
ASP
V
224
R
-
ND1
HIS
A
555
L
H
CG
ASP
V
224
R
-
ND1
HIS
A
555
L
H
OD2
ASP
V
224
R
-
ND1
HIS
A
555
L
H
HB2
ASP
V
224
R
-
ND1
HIS
A
555
L
H
HB3
ASP
V
224
R
-
ND1
HIS
A
555
L
H
H
TYR
V
225
R
-
CE1
HIS
A
555
L
H
HA
LEU
V
131
R
-
CE1
HIS
A
555
L
H
HD11
LEU
V
131
R
-
CE1
HIS
A
555
L
H
HD13
LEU
V
131
R
-
CE1
HIS
A
555
L
H
HG2
GLU
V
134
R
-
CE1
HIS
A
555
L
H
CA
ALA
V
221
R
-
CE1
HIS
A
555
L
H
C
ALA
V
221
R
-
CE1
HIS
A
555
L
H
CB
ALA
V
221
R
-
CE1
HIS
A
555
L
H
O
ALA
V
221
R
-
CE1
HIS
A
555
L
H
HA
ALA
V
221
R
-
CE1
HIS
A
555
L
H
HB1
ALA
V
221
R
-
CE1
HIS
A
555
L
H
CA
ASP
V
224
R
-
CE1
HIS
A
555
L
H
C
ASP
V
224
R
-
CE1
HIS
A
555
L
H
CB
ASP
V
224
R
-
CE1
HIS
A
555
L
H
CG
ASP
V
224
R
-
CE1
HIS
A
555
L
H
OD2
ASP
V
224
R
-
CE1
HIS
A
555
L
H
H
ASP
V
224
R
-
CE1
HIS
A
555
L
H
HB2
ASP
V
224
R
-
CE1
HIS
A
555
L
H
HB3
ASP
V
224
R
-
CE1
HIS
A
555
L
H
N
TYR
V
225
R
-
CE1
HIS
A
555
L
H
CA
TYR
V
225
R
-
CE1
HIS
A
555
L
H
H
TYR
V
225
R
-
CE1
HIS
A
555
L
H
HB2
TYR
V
225
R
-
NE2
HIS
A
555
L
H
CA
ALA
V
221
R
-
NE2
HIS
A
555
L
H
C
ALA
V
221
R
-
NE2
HIS
A
555
L
H
O
ALA
V
221
R
-
NE2
HIS
A
555
L
H
HA
ALA
V
221
R
-
NE2
HIS
A
555
L
H
N
ASP
V
224
R
-
NE2
HIS
A
555
L
H
CA
ASP
V
224
R
-
NE2
HIS
A
555
L
H
C
ASP
V
224
R
-
NE2
HIS
A
555
L
H
CB
ASP
V
224
R
-
NE2
HIS
A
555
L
H
O
ASP
V
224
R
-
NE2
HIS
A
555
L
H
CG
ASP
V
224
R
-
NE2
HIS
A
555
L
H
OD2
ASP
V
224
R
-
NE2
HIS
A
555
L
H
H
ASP
V
224
R
-
NE2
HIS
A
555
L
H
HA
ASP
V
224
R
-
NE2
HIS
A
555
L
H
HB2
ASP
V
224
R
-
NE2
HIS
A
555
L
H
HB3
ASP
V
224
R
-
NE2
HIS
A
555
L
H
N
TYR
V
225
R
-
NE2
HIS
A
555
L
H
CA
TYR
V
225
R
-
NE2
HIS
A
555
L
H
CB
TYR
V
225
R
-
NE2
HIS
A
555
L
H
H
TYR
V
225
R
-
NE2
HIS
A
555
L
H
HA
TYR
V
225
R
-
NE2
HIS
A
555
L
H
HB2
TYR
V
225
R
-
NE2
HIS
A
555
L
H
HB3
TYR
V
225
R
-
NE2
HIS
A
555
L
H
HE21
GLN
V
228